1 |
Needles with basal sheath deciduous and basal scalelike leaves not decurrent, cross section with 1 vascular bundle; umbo terminal or dorsal. |
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(2) |
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Needles with basal sheath persistent and basal scalelike leaves decurrent, cross section with 2 vascular bundles; umbo dorsal. |
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(16) |
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2 (1) |
Umbo dorsal; needles 3-5 per bundle. |
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(3) |
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Umbo terminal; needles (2-)5 per bundle. |
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(5) |
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3 (2) |
Needles 4 or 5 per bundle; bark dark brown, flaking |
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25 Pinus squamata |
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Needles 3 per bundle; bark white, irregularly flaking. |
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(4) |
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4 (3) |
Seed cones 12-20 cm; seeds cylindric, ca. 2.5 cm |
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26 Pinus gerardiana |
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Seed cones 5-7 cm; seeds ovoid, ca. 1 cm |
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27 Pinus bungeana |
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5 (2) |
Seeds wingless or only shortly winged. |
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(6) |
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Seeds long winged. |
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(10) |
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6 (5) |
Seed cones dehiscent at maturity; branchlets glabrous. |
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(7) |
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Seed cones indehiscent or imperfectly dehiscent; branchlets densely pubescent. |
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(8) |
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7 (6) |
Seeds wingless or ridged along abaxial margin; seed coat thick; apophyses not or slightly reflexed.at margin |
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31 Pinus armandi |
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Seeds with a rudimentary wing 2-7 mm; seed coat thin; apophyses obviously reflexed at margin |
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32 Pinus fenzeliana |
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8 (6) |
Needles slender, 4-8 cm × 0.5-1 mm; resin canals 2, marginal; shrubs |
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28 Pinus pumila |
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Needles stout, 7-12 cm × 1-1.5 mm; resin canals 3, median; trees. |
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(9) |
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9 (8) |
First-year branchlets with red-brown hairs; needles with 6-8 blue-gray stomatal lines along each abaxial surface; seed cones 9-14 cm; seed scales acuminate at apex, recurved |
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29 Pinus koraiensis |
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First-year branchlets with yellow-brown hairs; needles with 3-5 gray-white stomatal lines along each abaxial surface; seed cones 5-8 cm; seed scales rounded at apex, incurved |
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30 Pinus sibirica |
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10 (5) |
Needles 7-20 cm; seed cones cylindric to narrowly cylindric, 7-25 cm. |
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(11) |
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Needles less than 8 cm; seed cones ovoid, ovoid-ellipsoid, or ellipsoid-cylindric, usually less than 10 cm. |
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(13) |
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11 (10) |
Needles not pendulous, 6-14 cm; seed cones 7-20 cm |
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38 Pinus strobus |
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Needles pendulous, (6-)11-24 cm; seed cones 10-30 cm. |
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(12) |
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12 (11) |
Needles dark green adaxially; 1st-year branchlets green with faint bloom, glabrous; abaxial resin canal of needles always median |
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34 Pinus wallichiana |
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Needles grass green adaxially, conspicuously whitish bloomed on 2 abaxial surfaces; 1st-year branchlets conspicuously whitish bloomed, glandular pubescent; 1(or 2) abaxial resin canal(s) of needles always asymmetrically placed |
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33 Pinus bhutanica |
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13 (10) |
Needles less than 1 mm in diam.; seed cones sessile or shortly pedunculate. |
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(14) |
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Needles 1-1.5 mm in diam.; seed cones distinctly pedunculate. |
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(15) |
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14 (13) |
Branchlets densely pubescent; seed cones sessile; seeds as long as broad wing; needles 3.5-5.5 cm |
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39 Pinus parviflora |
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Branchlets glabrous or puberulent; seed cones shortly pedunculate; seeds ca. 1/2 as long as narrow wing; needles 4-8 cm |
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35 Pinus morrisonicola |
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15 (13) |
Branchlets densely pubescent; resin canals 3, median |
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36 Pinus wangii |
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Branchlets glabrous, rarely puberulent; resin canals 2, marginal, sometimes also 1 median |
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37 Pinus kwangtungensis |
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16 (1) |
Wing adnate to seed; needles 3 per bundle; apophyses distinctly protruded, with transverse ridges and spiny umbo |
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1 Pinus roxburghii |
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Wing articulated to seed; needles 2-5 per bundle; apophyses and umbo of different types. |
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(17) |
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17 (16) |
Needles (2 or)3(-5) per bundle. |
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(18) |
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Needles 2(or 3) per bundle. |
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(28) |
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18 (17) |
Trunk and branches usually with adventitious sprouts; needles mostly to 20 cm. |
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(19) |
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Trunk and branches usually without adventitious sprouts; needles mostly 15-45 cm. |
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(21) |
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19 (18) |
Needles 2(or 3) per bundle, ca. 1 mm in diam.; winter buds 0.5-0.7(-1) cm in diam.; bark plates with evident resin pockets; adaxial surface of seed scales lacking contrasting border distally |
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11 Pinus echinata |
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Needles 3(-5) per bundle, 1-1.5(-2) mm in diam.; winter buds 1-1.5(-2) cm in diam.; bark plates without evident resin pockets; adaxial surface of seed scales with dark red-brown border distally. |
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(20) |
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20 (19) |
Needles (12-)15-21 cm, slightly twisted; branchlets initially orange or yellow-orange, often glaucous; seed cones late dehiscent after maturity |
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12 Pinus serotina |
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Needles 5-10(-15) cm, twisted; branchlets initially orange-brown, not glaucous; seed cones dehiscent at maturity |
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13 Pinus rigida |
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21 (18) |
Needles slender, 10-30 cm × 0.7-1.2 mm (except P. yunnanensis var. pygmaea with stout needles 5-13 cm). |
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(22) |
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Needles ± stout, 12-45 cm × 1.5-2 mm. |
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(24) |
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22 (21) |
Umbo blunt or shortly mucronate in apical part of cone; needles 2(or 3) per bundle, 12-20 cm, slightly twisted |
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2 Pinus massoniana |
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Umbo shortly mucronate; needles (2 or)3 per bundle, 10-30 cm, not twisted. |
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(23) |
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23 (22) |
Needles slender and pliant, less than 1 mm in diam.; resin canals marginal; apophyses ± pyramidal, distinctly cross keeled |
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3 Pinus kesiya |
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Needles relatively stout and stiff (slender and pliant in plants growing in dry, hot valleys), 1-1.2 mm in diam.; resin canals marginal and median; apophyses swollen, usually not pyramidal, not or slightly cross keeled |
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4 Pinus yunnanensis |
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24 (21) |
Winter buds 3-4 cm, silvery white; seed cones 15-25 cm |
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14 Pinus palustris |
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Winter buds to 2 cm, reddish, chestnut, orange, or silvery brown; seed cones usually 5-15 cm. |
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(25) |
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25 (24) |
Seed cones pedunculate. |
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(26) |
+ |
Seed cones sessile or subsessile. |
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(27) |
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26 (25) |
Seed cones (7-)9-18(-20) cm, all deciduous; umbo central, depressed-pyramidal, with a stout, short prickle; needles 2 or 3 per bundle |
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15 Pinus elliottii |
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Seed cones 5-10(-12) cm, often a few basal cones persistent; umbo slightly projecting, ending in a straight, minute prickle; needles (2 or)3(-5) per bundle |
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16 Pinus caribaea |
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27 (25) |
Branchlets ca. 1 cm in diam.; terminal bud lanceolate-cylindric, mostly less than 1 cm wide, slightly resinous; seed cones mostly dull yellow-brown, narrowly ovoid when open |
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17 Pinus taeda |
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Branchlets 1-2 cm in diam.; terminal bud ovoid, ca. 1 cm wide, very resinous; seed cones mostly reddish brown, broadly ovoid when open |
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18 Pinus ponderosa |
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28 (17) |
Needles mostly 20-30 cm or more. |
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(29) |
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Needles mostly less than 20 cm. |
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(33) |
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29 (28) |
Umbo blunt or with sunken, short mucro. |
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(30) |
+ |
Umbo ± spiny or pointed at apex. |
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(31) |
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30 (29) |
Needle with 2 resin canals; seed cones red-brown at maturity |
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6 Pinus latteri |
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Needle with up to 9 resin canals; seed cones pale brown at maturity |
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19 Pinus tropicalis |
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31 (29) |
Seed cones borne on stalks to 3 cm, chocolate brown; apophyses lustrous |
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15 Pinus elliottii |
+ |
Seed cones sessile or subsessile, brown, yellow-, or red-brown; apophyses rarely lustrous. |
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(32) |
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32 (31) |
Branchlets ca. 1 cm in diam.; terminal bud lanceolate-cylindric, usually less than 1 cm wide, slightly resinous; seed cones usually dull yellow-brown, narrowly ovoid when open |
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17 Pinus taeda |
+ |
Branchlets 1-2 cm in diam.; terminal bud ovoid, ca. 1 cm wide, very resinous; seed cones usually reddish brown, broadly ovoid when open |
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18 Pinus ponderosa |
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33 (28) |
Seed cones 9-18 cm |
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20 Pinus pinaster |
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Seed cones usually not more than 8 cm. |
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(34) |
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34 (33) |
Needles slender and pliant, ca. 1 mm in diam.; apophyses relatively thin, flat or slightly raised. |
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(35) |
+ |
Needles stout, 1-2 mm in diam.; apophyses ± swollen or pyramidal. |
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(37) |
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35 (34) |
Trunk and branches usually with adventitious sprouts; umbo with a stout, sharp prickle |
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11 Pinus echinata |
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Trunk and branches usually without adventitious sprouts; umbo flat or blunt. |
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(36) |
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36 (35) |
Needles 12-20 cm, slightly twisted |
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2 Pinus massoniana |
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Needles 5-12 cm, straight 7. P. densiflora |
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(34) |
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37 (34) |
Needles shorter, longest ones not more than 8 cm, strongly twisted. |
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(38) |
+ |
Needles (5-)9-16 cm, straight or rarely slightly twisted. |
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(40) |
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38 (37) |
Seed cones strongly asymmetric, curved forward on branches; needles 2-5 cm |
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23 Pinus banksiana |
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Seed cones symmetric or nearly so; needles usually longer. |
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(39) |
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39 (38) |
Bark on distal sections of trunk orange, plated; needles blue-, gray-, or yellow-green, stomatal lines conspicuous; branchlets initially dull green or orange-brown, not glaucous; adaxial surface of seed scales lacking contrasting border distally |
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10 Pinus sylvestris |
+ |
Bark on distal sections of trunk reddish, scaly; needles deep to pale yellow-green, stomatal lines inconspicuous; branchlets initially red or purple tinged, often glaucous; adaxial surface of seed scales with strong purple-red or -brown border distally |
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24 Pinus virginiana |
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40 (37) |
Seed cones deciduous. |
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(41) |
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Seed cones persistent. |
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(42) |
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41 (40) |
Winter buds silvery white, cylindric-ellipsoid or cylindric |
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21 Pinus thunbergii |
+ |
Winter buds brown, ovoid or oblong-ovoid |
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22 Pinus nigra |
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42 (40) |
Needle with resin canals median, rarely also marginal; needles 4.5-17 cm; seed cones 3-6 cm |
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8 Pinus taiwanensis |
+ |
Needle with resin canals marginal, occasionally 1 or 2 median; needles 6-15 cm; seed cones 2.5-9 cm. |
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(43) |
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43 (42) |
Bark initially reddish brown, becoming dark grayish brown; 1st-year branchlets shiny yellowish brown; needle sheaths initially 0.5-1 cm; resin canals 3 or 4; pollen cones 1-1.8 cm; seed cones shiny chocolate brown at maturity |
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5 Pinus densata |
+ |
Bark dull grayish brown or dark gray; 1st-year branchlets light grayish brown, light brown, or yellowish brown; needle sheaths initially 1-2 cm; resin canals 5-9; pollen cones 0.5-0.9 cm; seed cones yellowish brown to brown at maturity |
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9 Pinus tabuliformis |