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FOC | Family List | FOC Vol. 12 | Rhamnaceae

6. Rhamnella Miquel, Prolus. Fl. Jap. 218; Ann. Mus. Bot. Lugduno-Batavi. 3: 30. 1867.

猫乳属 mao ru shu

Chaydaia Pitard.

Shrubs or small trees, rarely scandent, evergreen to mostly deciduous. Leaves alternate, shortly petiolate; stipules narrowly triangular or lanceolate, often persistent; leaf blade papery or submembranous, pinnately veined, margin serrulate. Flowers yellow-green, small, bisexual, 5-merous, pedicellate, in shortly pedunculate axillary cymes, or fascicled in axils of leaves. Calyx tube cup-shaped; lobes triangular, midvein adaxially keeled and with centrally inserted beaklike protrusion. Petals ovate-spatulate or orbicular-spatulate, ± distinctly cucullate, enfolding stamens. Stamens dorsifixed; filaments free from claw at base, lanceolate-linear. Disk pentagonous, thin, lining calyx tube, nectariferous. Ovary superior, base slightly immersed in disk, 1- or 2-loculed or imperfectly 1-loculed, with 2 ovules per locule; styles terminally 2-fid. Drupe yellowish to orange, turning black or purple-black when ripe, cylindric-ellipsoidal base with persistent remnants of calyx tube, with rudimentary style at apex, 1- or 2-loculed, with 1 or 2 seeds.

Eight species: China, Japan, Korea; eight species (five endemic) in China.

1 Evergreen shrubs or small trees, rarely scandent   1 R. rubrinervis
+ Deciduous shrubs or small trees, never scandent   (2)
2 (1) Young branches, petioles, and abaxial leaf surface pubescent   (3)
+ Young branches, petioles, and abaxial leaf surface glabrous or subglabrous   (4)
3 (2) Leaf blade obovate-oblong or obovate-elliptic, rarely obovate, apex caudate-acuminate or long acuminate, abaxially pubescent or only on veins, lateral veins 5-11(-13) pairs.   2 R. franguloides
+ Leaf blade subelliptic or oblong, apex shortly acuminate or acuminate, abaxially tomentose or pubescent, lateral veins 5-8 pairs.   3 R. julianae
4 (2) Leaf blade 7-15 cm, obovate-elliptic or obovate-oblong, apex caudate-acuminate, glabrous or abaxially pilose on veins, lateral veins 7-10 pairs.   4 R. caudata
+ Leaf blade usually shorter than 7 cm, ovate, ovate-elliptic, or lanceolate-elliptic, apex acute or acuminate, lateral veins 4-8 pairs   (5)
5 (4) Leaf lateral veins 4 or 5 pairs, margin subentire or inconspicuously serrulate   (6)
+ Leaf lateral veins 5-8 pairs, margin conspicuously serrulate   (7)
6 (5) Leaf blade ovate or ovate-elliptic, apex shortly acuminate, rarely acute, margin entire at lower half, inconspicuously serrulate at upper half.   5 R. wilsonii
+ Leaf blade elliptic or lanceolate-elliptic, broadest at middle, apex acute, margin inconspicuously serrulate.   6 R. gilgitica
7 (5) Leaf blade oblong, both surfaces glabrous, margin entire at lower 1/3, lateral veins 5-7 per side.   7 R. forrestii
+ Leaf blade narrowly elliptic, adaxially pilose on veins, margin serrulate, lateral veins 6-8 pairs.   8 R. martini

Lower Taxa


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