1 |
Flowers bisexual. |
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(2) |
+ |
Flowers unisexual (shrubs dioecious). |
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(31) |
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2 (1) |
Branchlets with spines, prickles, or both. |
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(3) |
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Branchlets with neither spines nor prickles. |
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(10) |
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3 (2) |
Flowers 4-20 in racemes more than 2 cm |
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1 R. horridum |
+ |
Flowers solitary, 2 or 3 in clusters, or in racemes less than 2 cm. |
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(4) |
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4 (3) |
Style and calyx tube adaxially villous |
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2 R. reclinatum |
+ |
Style and calyx tube adaxially glabrous. |
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(5) |
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5 (4) |
Fruit oblong, 2-2.5 cm |
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3 R. stenocarpum |
+ |
Fruit globose, rarely ellipsoid, 0.8-1.5 cm. |
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(6) |
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6 (5) |
Spines at nodes slender, basally less than 1 mm in diam. |
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(7) |
+ |
Spines at nodes robust, basally more than 1 mm in diam. |
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(8) |
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7 (6) |
Leaf blade glabrous except at vein axils; style divided for ca. 1/2 its length |
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4 R. aciculare |
+ |
Leaf blade pubescent, some hairs glandular; style entire or only stigma 2-lobed |
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5 R. fuyunense |
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8 (6) |
Calyx tube cylindric, 7-10 mm |
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6 R. formosanum |
+ |
Calyx tube campanulate, to 6 mm. |
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(9) |
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9 (8) |
Style 2-lobed at apex; anthers not exserted |
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7 R. burejense |
+ |
Style divided for ca. 1/2 its length; anthers exserted |
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8 R. alpestre |
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10 (2) |
Flowers solitary or 2 per fascicle |
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9 R. ambiguum |
+ |
Flowers in racemes. |
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(11) |
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11 (10) |
Shrubs low, creeping, or procumbent, to 40 cm tall, rarely semierect to 1 m tall. |
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(12) |
+ |
Shrubs erect, usually more than 1 m tall. |
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(14) |
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12 (11) |
Leaf blade with scattered, sessile, yellow glands abaxially |
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10 R. procumbens |
+ |
Leaf blade without scattered, sessile, yellow glands abaxially, sometimes with glandular hairs. |
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(13) |
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13 (12) |
Racemes pendent, lax, (3-)5-7-flowered; calyx purple to red |
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11 R. triste |
+ |
Racemes erect, dense, 5-15-flowered; calyx yellowish white |
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12 R. palczewskii |
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14 (11) |
Racemes short, 2- or 3-flowered |
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13 R. fargesii |
+ |
Racemes (2-)4-30(-40) cm, (4-)8-40(-50)-flowered. |
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(15) |
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15 (14) |
Calyx tube cylindric, 1.2-1.5 cm |
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14 R. odoratum |
+ |
Calyx tube campanulate or cupular, or if cylindric then less than 0.7 cm. |
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(16) |
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16 (15) |
Leaf blade with scattered, sessile, yellow glands abaxially. |
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(17) |
+ |
Leaf blade without scattered, sessile, yellow glands abaxially, sometimes with glandular hairs. |
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(18) |
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17 (16) |
Calyx and ovary with scattered, sessile, yellow glands |
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15 R. nigrum |
+ |
Calyx and ovary without scattered, sessile, yellow glands |
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16 R. americanum |
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18 (16) |
Calyx lobes reflexed at or soon after anthesis. |
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(19) |
+ |
Calyx lobes erect or incurved at anthesis. |
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(23) |
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19 (18) |
Racemes lax, rachis clearly visible at anthesis. |
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(20) |
+ |
Racemes dense, rachis almost hidden at anthesis. |
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(21) |
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20 (19) |
Petiole 6-8 cm, leaf lobes acuminate at apex; calyx lobes not ciliate; fruit red, 0.8-1.2 cm |
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17 R. griffithii |
+ |
Petiole 3-5 cm, leaf lobes obtuse to acute at apex; calyx lobes ciliate; fruit purple-black, 0.5-0.7 cm |
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18 R. altissimum |
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21 (19) |
Racemes spreading then pendulous, fewer than 30-flowered; calyx purple, without warts at base of calyx tube adaxially |
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19 R. soulieanum |
+ |
Racemes erect then pendulous, usually more than 30-flowered; calyx green to yellow, with 5 warts at base of calyx tube adaxially. |
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(22) |
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22 (21) |
Leaf lobes coarsely sharply serrate or doubly serrate at margin, acute or shortly acu-minate at apex; warts on adaxial surface of calyx tube not joined by a raised ring |
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20 R. mandshuricum |
+ |
Leaf lobes coarsely crenate-serrate or doubly serrate at margin, obtuse or acute at apex; warts on adaxial surface of calyx tube joined by a raised ring |
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21 R. multiflorum |
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23 (18) |
Calyx lobes ciliate or dentate at margin. |
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(24) |
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Calyx lobes neither ciliate nor dentate at margin, at most erose. |
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(27) |
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24 (23) |
Racemes lax, rachis clearly visible at anthesis; calyx tube pelviform, 1-1.5 mm |
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22 R. pubescens |
+ |
Racemes dense, rachis almost hidden at anthesis; calyx tube campanulate to shortly cylindric, 1.5-3.5 mm. |
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(25) |
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25 (24) |
Petals not ciliate at margin; leaf blade to 12 × 15 cm, petiole 5-8 cm |
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23 R. latifolium |
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Petals ciliate at margin; leaf blade to 10 × 10 cm, petiole 2.5-5 cm. |
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(26) |
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26 (25) |
Leaf blade and raceme rachis stalked glandular; petals subspatulate to flabellate; stamens inserted level with petals |
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24 R. himalense |
+ |
Leaf blade and raceme rachis not stalked glandular; petals narrowly cuneate to linear; stamens inserted below petals |
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25 R. meyeri |
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27 (23) |
Racemes initially erect, 2-6 cm, 5-15-flowered; calyx tube pelviform or shallowly cupular, 1-2 mm. |
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(28) |
+ |
Racemes nodding to pendulous, 5-40 cm, 9-50-flowered; calyx tube campanulate to shortly cylindric, 2-6 mm. |
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(29) |
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28 (27) |
Calyx green to greenish brown; petals purplish; raceme rachis and pedicels sparsely shortly glandular hairy |
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26 R. rubrum |
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Calyx yellowish white; petals yellowish; raceme rachis and pedicels pubescent at anthesis, glabrous in fruit |
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12 R. palczewskii |
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29 (27) |
Racemes lax, 15-40-flowered; pedicels 3-10(-15) mm; calyx tube 4-6 mm; stamens exserted |
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27 R. longeracemosum |
+ |
Racemes dense, 5-30(-50)-flowered; pedicels absent or very short; calyx tube 2-4 mm; stamens included. |
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(30) |
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30 (29) |
Buds glabrous, apex obtuse; leaf blade adaxially sparsely coarsely glandular hairy; calyx lobes not incurved, apex obtuse; stamens inserted level with petals |
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28 R. moupinense |
+ |
Buds puberulent, apex acute; leaf blade adaxially densely pubescent; calyx lobes incurved, apex acute; stamens inserted below petals |
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29 R. setchuense |
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31 (1) |
Shrubs deciduous, though in leaf for most of year; flowers in subsessile umbels or fascicles, rarely solitary |
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30 R. fasciculatum |
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Shrubs deciduous, semievergreen, or evergreen; flowers in racemes. |
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(32) |
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32 (31) |
Shrubs evergreen, rarely semievergreen, unarmed; leaf blade margin entire, rarely 3-lobed near apex. |
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(33) |
+ |
Shrubs deciduous, unarmed or with 2 small prickles at each node; leaf blade margin lobed. |
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(37) |
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33 (32) |
Branchlets and petioles glandular bristly; petioles 3-8 mm. |
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(34) |
+ |
Branchlets glabrous; petioles 0.5-4 cm, glabrous or sparsely glandular hairy. |
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(35) |
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34 (33) |
Leaf blade narrowly elliptic to sublanceolate, ca. 3 × as long as wide; calyx purplish green, glabrous |
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31 R. tianquanense |
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Leaf blade elliptic to obovate-elliptic, ca. 2 × as long as wide; calyx greenish white, stalked glandular |
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32 R. henryi |
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35 (33) |
Leaf blade orbicular to reniform-orbicular, about as long as wide; calyx purple to purplish green |
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33 R. hunanense |
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Leaf blade obovate to elliptic, longer than wide; calyx greenish white or yellowish green. |
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(36) |
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36 (35) |
Leaf blade 2-5 × 1.5-3 cm; male racemes erect; fruit 0.8-1.1 × 0.6-0.8 cm |
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34 R. davidii |
+ |
Leaf blade 5-10 × 2.5-4.5 cm; male racemes pendulous; fruit 1.5-2 × 0.7-1 cm |
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35 R. laurifolium |
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37 (32) |
Nodes each with 2 small spines; internodes minutely prickly or unarmed. |
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(38) |
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Nodes and internodes unarmed. |
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(42) |
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38 (37) |
Petioles pubescent to glandular hairy, rarely glabrescent; leaf blade margin 3-5-lobed. |
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(39) |
+ |
Petioles glabrous, rarely puberulent; leaf blade margin 3-lobed. |
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(40) |
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39 (38) |
Female racemes up to 6-flowered; calyx tube 2-3 mm; fruit pubescent then glabrescent |
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36 R. giraldii |
+ |
Female racemes at least 8-flowered; calyx tube 1.5-2 mm; fruit glabrous |
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37 R. pulchellum |
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40 (38) |
Shrubs low, to 1 m tall; leaf blade puberulent, remaining so at margin and sometimes abaxially |
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38 R. saxatile |
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Shrubs erect, to 3 m tall; leaf blade glabrous. |
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(41) |
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41 (40) |
Petioles 0.5-1 cm; leaf blade 1-1.5 cm; male racemes 7-11-flowered |
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39 R. glabrifolium |
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Petioles 1-2 cm; leaf blade 1.5-3.5 cm; male racemes 10-15-flowered |
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40 R. diacanthum |
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42 (37) |
Terminal leaf lobe shorter than or rarely equaling lateral ones, apex obtuse, rarely acute. |
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(43) |
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Terminal leaf lobe equaling or longer than lateral ones, apex acute, acuminate, or caudate. |
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(46) |
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43 (42) |
Leaf blade glabrous or margin ciliate, rarely viscid glandular; ovary and fruit glabrous |
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41 R. heterotrichum |
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Leaf blade pubescent, or viscid or stalked glandular, or both; ovary and young fruit pubescent and glandular pubescent. |
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(44) |
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44 (43) |
Calyx green, sometimes tinged reddish brown |
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42 R. kialanum |
+ |
Calyx purple or purplish brown. |
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(45) |
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45 (44) |
Leaf blade orbicular to reniform-orbicular, pubescent and glandular, lobes coarsely obtusely serrate at margin, obtuse at apex; calyx pubescent and glandular |
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43 R. orientale |
+ |
Leaf blade broadly ovate, glandular but not pubescent, lobes sharply serrate at margin, obtuse to acute at apex; calyx glabrous, rarely puberulent |
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44 R. xizangense |
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46 (42) |
Leaf blade to 2 cm, terminal lobe slightly longer than lateral ones. |
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(47) |
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Leaf blade more than 2 cm, terminal lobe usually much longer than lateral ones. |
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(49) |
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47 (46) |
Calyx pubescent; fruit black or reddish black |
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45 R. pseudofasciculatum |
+ |
Calyx glabrous; fruit red. |
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(48) |
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48 (47) |
Calyx purple; leaf blade glabrous, rarely sparsely glandular hairy adaxially; inflorescences glabrous |
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46 R. humile |
+ |
Calyx green or yellow-green; leaf blade and inflorescences minutely pubescent and stalked glandular |
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47 R. glabricalycinum |
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49 (46) |
Calyx hairy or glandular or both. |
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(50) |
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Calyx glabrous. |
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(55) |
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50 (49) |
Fruit black. |
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(51) |
+ |
Fruit yellowish green, red, or reddish brown. |
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(52) |
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51 (50) |
Calyx green, sometimes tinged reddish brown, tube cupular; fruit sparsely stalked glandular, sometimes hairy |
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48 R. vilmorinii |
+ |
Calyx dark red or purple, tube saucer-shaped, rarely shallowly cupular; fruit glabrous |
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49 R. rubrisepalum |
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52 (50) |
Fruit densely villous and stalked glandular. |
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(53) |
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Fruit glabrous, pubescent, or sparsely stalked glandular. |
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(54) |
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53 (52) |
Leaf blade 2.5-5 cm, margin acutely serrate or doubly serrate; male racemes lax, 6-7 cm, 10-15-flowered |
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50 R. franchetii |
+ |
Leaf blade 6-10 cm, margin obtusely dentate or doubly dentate; male racemes dense, 7-15 cm, 15-30-flowered or more |
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51 R. maximowiczii |
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54 (52) |
Leaf blade 5-9 cm, margin deeply doubly serrate, terminal lobe longer than lateral ones, apex acuminate; calyx reddish brown |
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52 R. takare |
+ |
Leaf blade 2-5 cm, margin irregularly crenate-dentate or doubly dentate, rarely subentire, terminal lobe subequaling lateral ones; calyx green, sometimes tinged reddish brown |
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42 R. kialanum |
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55 (49) |
Inflorescences stalked glandular, otherwise glabrous. |
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(56) |
+ |
Inflorescences hairy and stalked glandular. |
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(57) |
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56 (55) |
Leaf blade glabrous, rarely sparse ly stalked glandular, terminal lobe acute at apex; calyx green |
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53 R. komarovii |
+ |
Leaf blade coarsely appressed hairy adaxially, terminal lobe acuminate at apex; calyx yellowish brown |
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54 R. maximowiczianum |
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57 (55) |
Fruit black. |
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(58) |
+ |
Fruit red or dark purple. |
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(60) |
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58 (57) |
Terminal leaf lobe usually less than 2 × as long as lateral ones |
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55 R. luridum |
+ |
Terminal leaf lobe usually more than 2 × as long as lateral ones. |
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(59) |
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59 (58) |
Male racemes 1.5-2.5 cm; pedicels 1-2 mm; calyx green, sometimes slightly tinged reddish brown |
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+ |
desc needed |
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