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45. SANGUISORBA Linnaeus, Sp. Pl. 1: 116. 1753.
地榆属 di yu shu
Li Chaoluan (Li Chao-luang); Hiroshi Ikeda, Hideaki Ohba
Poterium Linnaeus.
Herbs perennial. Rootstock robust, bearing many fusiform, cylindric roots in lower part. Stipules sheathing, adnate to petiole, petiole sheathing and imbricate at base; leaf blade imparipinnate; leaflets serrate at margin. Inflorescences terminal on elongate scapes, densely capitate or spicate, bracteate and bracteolate. Flowers bisexual, rarely unisexual (when plants monoecious). Hypanthium with a constricted throat. Sepals 4(–7), imbricate, petaloid, purple, red, pink, or white, rarely greenish. Petals absent. Disk lining hypanthium. Stamens usually 4, rarely more, inserted in throat of hypanthium; filaments free, rarely partly connate; anthers didymous. Carpel 1(or 2), included in hypanthium; ovule pendulous; style terminal, filiform; stigma penicillate. Achene dry, included in hardened, muricate or winged hypanthium. Cotyledons planoconvex. x = 7.
About 30 species: Asia, Europe, North America; seven species (one endemic) in China.
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1 |
Inflorescences flowering from base upward. |
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(2) |
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Inflorescences flowering from apex downward. |
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(3) |
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2 (1) |
Leaflets of radical leaves usually truncate to subcordate at base; inflorescences usually nodding, thick; sepals white or pinkish; bracts longer than hypanthium in bud, densely hairy; filaments gradually dilated from base, broadest near middle, then gradually attenuate toward apex, narrower than anther, or sometimes little dilated. |
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6 S. alpina |
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Leaflets of radical leaves cordate or deeply so at base; inflorescences usually erect, long, slender; sepals white; bracts nearly equaling hypanthium in bud, abaxially sparsely hairy or subglabrous, margin ciliate; filaments compressed-dilated from middle to apex, broadest at apex, nearly as broad as anther. |
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7 S. stipulata |
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3 (1) |
Inflorescences capitate; sepals white. |
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(4) |
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Inflorescences ellipsoid, cylindric, or elongate; sepals purple, red, pink or white. |
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(5) |
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4 (3) |
Plants less than 35 cm tall; leaflets of radical leaves broadly ovate to suborbicular; flowers unisexual (females in center, males surrounding); style 0.5–1 × as long as sepals; fruiting hypanthium with 4 ribs. |
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4 S. filiformis |
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Plants to 85 cm tall; leaflets of radical leaves ovate, elliptic, or long elliptic; flowers bisexual; style equaling sepals; fruiting hypanthium with 4 winged ribs. |
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5 S. diandra |
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5 (3) |
Filaments filiform, subequaling sepals. |
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1 S. officinalis |
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Filaments compressed-dilated, 0.5–2 × as long as sepals. |
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(6) |
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6 (5) |
Leaflets of radical leaves linear-lanceolate, base subcordate or rounded to obliquely broadly cuneate, margin acutely incised serrate. |
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2 S. tenuifolia |
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Leaflets of radical leaves ovate, elliptical, or oblong, base cordate, margin coarsely obtusely serrate. |
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3 S. applanata |
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List of lower taxa
Related Links (opens in a new window) |
Treatments in Other Floras @ www.efloras.org
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