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60. Seseli Linnaeus, Sp. Pl. 1: 259. 1753.

西风芹属 xi feng qin shu

She Menglan (佘孟兰 Sheh Meng-lan); Michael G. Pimenov, Eugene V. Kljuykov, Mark F. Watson

Herbaceous, perennials, sometimes woody at base. Taproot conic, caudex woody, undivided or branched. Stem terete, rarely hollow. Leaf blade 1–3-pinnate or pinnately decompound, rarely ternately dissected or simple and undivided. Umbels compound; bracts few or absent; rays few to many; bracteoles few to many, lanceolate or linear, bases often connate, scarious or scarious-margined; umbellules few- to many-flowered; pedicels short or long, occasionally subsessile. Calyx teeth obsolete or minute. Petals white or yellow, suborbicular or oblong, emarginate, apex broadly or narrowly inflexed, abaxially often pubescent or hirsute. Stylopodium conic or depressed. Fruit ovoid or ellipsoid, moderately or slightly dorsally compressed, glabrous or variously hairy; ribs prominent, rounded or keeled, subequal, marginal ribs sometimes narrowly winged; vittae 1–2(–4) in each furrow, 2 (rarely 4–8) on commissure. Seed face plane. Carpophore 2-cleft to base.

About 80 species: Asia, Europe; 19 species (nine endemic) in China.

See the taxonomic comments under Eriocycla and Libanotis, and the alternative classification at the end of the Seseli account.

The following is a synopsis of an alternative classification of Seseli, including Libanotis and allied genera, as proposed by two of us (Pimenov and Kljuykov). Synonyms are included only where taxonomic concepts, as indicated through synonymy, differ from those used in the main accounts. Literature citations provided in the main accounts are not repeated.


1 Fruit vittae solitary in each furrow, 2 on the commissure.   (2)
+ Fruit vittae 2–5 in each furrow, 4–10 on the commissure.   (11)
       
2 (1) Caudex branched at apex, stems several.   (3)
+ Caudex undivided; stem solitary.   (7)
       
3 (2) Bracteoles glabrous; fruit papillose-pubescent; rays (2–)6–10.   (4)
+ Bracteoles puberulous; fruit pubescent or hirsute, but never papillose; rays 2–6(–10).   (5)
       
4 (3) Fruit with a white and membranous disk at base.   1 Seseli glabratum
+ Fruit without a white and membranous disk at base.   2 Seseli intramongolicum
       
5 (3) Umbels 1–2.5 cm across; bracteole bases connate.   3 Seseli aemulans
+ Umbels 2–6 cm across; bracteole bases free.   (6)
       
6 (5) Ultimate leaf segments 15–40(–80) × 0.7–1.5 mm; rays 3–4; flowers almost sessile.   14 Seseli togasii
+ Ultimate leaf segments 5–7 × 1–2 mm; rays 6–8(–10); flowers pedicellate.   15 Seseli junatovii
       
7 (2) Stem hollow.   4 Seseli nortonii
+ Stem solid, with pith.   (8)
       
8 (7) Plants densely white hispidulous; leaf blade 3-parted, not pinnate, ultimate segments 70–130 × 5–10 mm.   7 Seseli delavayi
+ Plants puberulent to almost glabrous; leaf blade 2–3-pinnate, ultimate segments 3–50 × 0.5–2 mm.   (9)
       
9 (8) Fruit oblong, 5–6.5 × 2–3 mm.   5 Seseli eriocephalum
+ Fruit ovoid, 2–4 × 0.8–1.5 mm.   (10)
       
10 (9) Ultimate leaf segments 5–12 × 0.5–1 mm; ribs slightly prominent, rounded.   6 Seseli valentinae
+ Ultimate leaf segments 20–50 × 0.5–2 mm; ribs prominent, keeled.   13 Seseli strictum
       
11 (1) Bracteoles fused to each other at base, sometimes, up to middle.   (12)
+ Bracteoles not fused at base.   (13)
       
12 (11) Leaf blade 2–4-ternately dissected, ultimate segments narrow-linear 7–65 × 0.5–3 mm.   8 Seseli yunnanense
+ Leaf blade trifoliolate or 2-ternately dissected, ultimate segments elliptic or lanceolate, 20–120 × 2–12 mm.   9 Seseli mairei
       
13 (11) Caudex branched.   (14)
+ Caudex unbranched.   (16)
       
14 (13) Flowers sessile, umbellules capitate.   10 Seseli sessiliflorum
+ Flowers pedicellate, umbels loose, not capitate.   (15)
       
15 (14) Leaf blade greenish pubescent; stems, rays and bracts scabrous; bracteoles 5–8; fruit minutely pubescent.   16 Seseli asperulum
+ Plant completely glabrous; bracteoles 8–10; fruit glabrous.   17 Seseli coronatum
       
16 (13) Stylopodium conic.   (17)
+ Stylopodium low-conic.   (18)
       
17 (16) Pedicels stout, 2–4 mm; leaf sheaths dark purple.   11 Seseli purpureovaginatum
+ Pedicels slender, short 0.5–1.5 mm; leaf sheaths green.   12 Seseli squarrulosum
       
18 (16) Fruit ovoid, apex narrow, densely pubescent; leaf blade 2-pinnate, ultimate segments ovate or rhombic.   18 Seseli sandbergiae
+ Fruit oblong glabrous; leaf blade 3-pinnate, ultimate segments ovate.   19 Seseli incisodentatum

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