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FOC | Family List | FOC Vol. 23 | Typhaceae

1. Sparganium Linnaeus, Sp. Pl. 2: 971. 1753.

黑三棱属 hei san leng shu

Herbs, perennial, aquatic or in marshes, with creeping rhizomes. Stems simple or branched, robust or slender. Leaves alternate, distichous, emergent or floating, sheathed at base. Flowers minute, densely crowded in globose heads on upper part of stems and branches; several male and female heads in panicle, raceme, or spike, male heads above female. Perianth segments of 3-6 chaffy elongate scales. Male flowers with 3 or more stamens; anthers basifixed, longitudinally dehiscent; filaments free or partly united. Female flowers bracteolate; ovary 1-loculed or rarely 2-loculed, narrow at base; styles simple or forked, with a unilateral stigma; ovule 1, pendulous. Fruit nutlike, indehiscent.

About 19 species: mainly in temperate and cool regions of the world, best represented in the N Hemisphere; 11 species (three endemic) in China.


1 Stems floating or slightly prostrate in water; leaves usually floating, flattened   (2)
+ Stems erect; leaves aerial, not flattened   (5)
       
2 (1) Stems stout, 1.3-1.5 m tall; leaves ca. 10 mm wide; male heads 8-10.   8 S. yunnanense
+ Stems slender, less than 70 cm tall; leaves 2-4 mm wide; male heads 1-4   (3)
       
3 (2) Styles absent or inconspicuous; fruit broadly obovoid.   11 S. hyperboreum
+ Styles conspicuous; fruit fusiform, elliptic, or lanceolate   (4)
       
4 (3) Stems floating, more than 20 cm; male heads 2-4.   9 S. angustifolium
+ Stems slightly prostate or rarely erect, less than 20 cm; male heads 1 or 2.   10 S. natans
       
5 (1) Inflorescences paniculate, with conspicuous lateral branches with male and female heads; ovary sessile   (6)
+ Inflorescences racemose or spicate, degenerated into 1 female head or completely degenerated; ovary with short stalk   (8)
       
6 (5) Panicles with 3-7 lateral branches; stigmas 3-4 mm.   1 S. stoloniferum
+ Panicles with 1(or 2) lateral branches; stigmas less than 2 mm   (7)
       
7 (6) Leaves 2-3 mm wide; main axis and lateral branch of panicles not zigzag; styles ca. 0.5 mm.   2 S. subglobosum
+ Leaves 4-5 mm wide; main axis and lateral branch of panicles usually zigzag; styles 1-1.5 mm.   3 S. limosum
       
8 (5) Inflorescences spicate; perianth segments lobate to parted.   7 S. confertum
+ Inflorescences racemose; perianth segments serrate-lobate to lobate   (9)
       
9 (8) Axis of racemes curved; ovary attenuate toward base and forming an inconspicuous stalk.   4 S. fallax
+ Axis of racemes not curved; ovary with a conspicuous stalk   (10)
       
10 (9) Male heads 4-8, distinctly separate from female ones; stigmas 1.5-1.8 mm.   5 S. emersum
+ Male heads 1-3, not separate from female ones; stigmas ca. 0.5 mm.   6 S. glomeratum

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