| 1 |
Inflorescences borne on erect, long, leafy shoots of current year arising from base of shrub, sometimes on short, lateral branchlets arising from branches of previous year. |
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(2) |
| + |
Inflorescences borne on short, lateral branchlets arising from (often arching) branches of previous year. |
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(30) |
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| 2 (1) |
Inflorescences paniculate, oblong or pyramidal. |
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1 S. salicifolia |
| + |
Inflorescences broadly compound corymbose. |
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(3) |
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| 3 (2) |
Inflorescences borne on erect, long, leafy shoots of current year. |
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(4) |
| + |
Inflorescences borne on short, lateral branchlets arising from branches of previous year. |
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(9) |
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| 4 (3) |
Inflorescences glabrous; flowers white; follicles erect. |
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7 S. fritschiana |
| + |
Inflorescences pubescent; flowers usually pink, rarely purple-red; follicles divergent. |
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(5) |
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| 5 (4) |
Leaf blade base truncate to rounded; petiole (2–)4–8 mm. |
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(6) |
| + |
Leaf blade base cuneate to broadly so; petiole 1–3 mm. |
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(7) |
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| 6 (5) |
Branchlets glabrous; leaf blades 3–7 cm, glabrous on both surfaces, margin crenate to doubly serrate; flowers bisexual, dark purple-red. |
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5 S. purpurea |
| + |
Branchlets pubescent or subglabrous; leaf blades 2–4 cm, slightly pubescent on veins on both surfaces or glabrous adaxially, margin sharply serrate or doubly serrate; flowers usually ± unisexual, pink, rarely white. |
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6 S. bella |
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| 7 (5) |
Leaf blades glabrous on both surfaces; inflorescences to 5 × 5.5 cm. |
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4 S. hayatana |
| + |
Leaf blades pubescent on both surfaces or only on veins; inflorescences 6–12 × 4–14 cm. |
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(8) |
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| 8 (7) |
Branchlets glabrous or pubescent when young; leaf blades doubly incised serrate at margin, rarely singly serrate, apex acute to acuminate. |
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2 S. japonica |
| + |
Branchlets densely yellow pubescent when young; leaf blades doubly serrate at margin, apex acute. |
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3 S. formosana |
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| 9 (3) |
Buds obtuse at apex, with several exterior scales. |
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(10) |
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Buds acute to acuminate at apex, with 2 exterior scales. |
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(19) |
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| 10 (9) |
Stamens 2–3 × as long as petals. |
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8 S. miyabei |
| + |
Stamens shorter than to nearly equaling petals. |
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(11) |
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| 11 (10) |
Leaf blade margin with a few teeth above middle or near apex. |
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(12) |
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Leaf blade margin entire, rarely with a few teeth apically. |
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(15) |
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| 12 (11) |
Follicles glabrous or hairy only along adaxial suture. |
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11 S. sargentiana |
| + |
Follicles hairy. |
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(13) |
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| 13 (12) |
Branchlets angled; leaf blades tomentose abaxially; pedicels 1–2 mm. |
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10 S. schochiana |
| + |
Branchlets terete; leaf blades densely thinly villous abaxially; pedicels 4–8 mm. |
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(14) |
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| 14 (13) |
Bracts 2–4 mm; follicles thinly villous. |
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9 S. henryi |
| + |
Bracts 4–6 mm; follicles densely pubescent. |
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12 S. wilsonii |
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| 15 (11) |
Leaf blades glabrous on both surfaces or thinly villous along margin; branchlets angled. |
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16 S. schneideriana |
| + |
Leaf blades softly hairy abaxially or on both surfaces; branchlets terete. |
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(16) |
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| 16 (15) |
Follicles glabrous. |
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15 S. veitchii |
| + |
Follicles softly hairy. |
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(17) |
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| 17 (16) |
Inflorescences glabrous; leaf blades glabrous or sparsely pubescent only on veins abaxially. |
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14 S. ovalis |
| + |
Inflorescences and both surfaces of leaf blades softly hairy. |
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(18) |
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| 18 (17) |
Inflorescences sparsely villous; leaf blades 10–30 × 5–10 mm, villous on both surfaces, adaxially without papillae, margin entire or apically with a few teeth. |
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12 S. wilsonii |
| + |
Inflorescences densely and thinly pubescent; leaf blades 6–11 × 4–7 mm, thinly pubescent on both surfaces when young, later glabrescent, adaxially with papillae, margin entire. |
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13 S. teretiuscula |
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| 19 (9) |
Leaf blade margin entire or with a few teeth above middle. |
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(20) |
| + |
Leaf blade margin singly or doubly serrate, sometimes incised. |
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(23) |
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| 20 (19) |
Branchlets cylindric or slightly angled; inflorescences glabrous. |
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28 S. uratensis |
| + |
Branchlets angled; inflorescences puberulous to pubescent. |
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(21) |
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| 21 (20) |
Flowers red; leaf blades 0.8–1.2 cm, sometimes lobed apically; follicles usually glabrous, rarely puberulous on adaxial suture. |
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25 S. arcuata |
| + |
Flowers white; leaf blades 1–3 cm, not lobed apically; follicles pubescent. |
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(22) |
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| 22 (21) |
Leaf blades ovate or obovate to obovate-lanceolate, 1–2 cm, pubescent abaxially, margin obtusely 3–5-dentate above middle. |
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26 S. canescens |
| + |
Leaf blades oblong to ovate- or obovate-oblong, 1.5–3 cm, glabrous on both surfaces, margin often entire. |
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27 S. trichocarpa |
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| 23 (19) |
Buds small, shorter than petioles. |
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(24) |
| + |
Buds usually large, nearly equaling or longer than petioles, very rarely shorter. |
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(27) |
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| 24 (23) |
Shrubs to 3 m tall; leaf blades abaxially slightly hairy on veins. |
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(25) |
| + |
Shrubs to 0.5 m tall; leaf blades abaxially glabrous. |
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(26) |
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| 25 (24) |
Leaf blades elliptic to elliptic-ovate, margin sharply singly serrate; inflorescences and hypanthium abaxially softly hairy. |
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17 S. teniana |
| + |
Leaf blades ovate to elliptic-ovate, margin obtusely singly or doubly serrate; inflorescences and hypanthium abaxially glabrous. |
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18 S. hingshanensis |
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| 26 (24) |
Leaf blades ovate or rhomboid-ovate, margin sharply singly serrate above middle; flowers white; ovary basally slightly pubescent. |
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19 S. morrisonicola |
| + |
Leaf blades ovate, margin doubly serrate; flowers pink; ovary glabrous. |
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20 S. kweichowensis |
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| 27 (23) |
Inflorescences and follicles subglabrous or pilose. |
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24 S. longigemmis |
| + |
Inflorescences and follicles densely softly hairy. |
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(28) |
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| 28 (27) |
Buds glabrous; leaf blades abaxially pubescent. |
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23 S. rosthornii |
| + |
Buds densely tomentose-villous; leaf blades abaxially sericeous-villous, sometimes glabrescent. |
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(29) |
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| 29 (28) |
Petioles 3–5 mm; leaf blades ovate or ovate-lanceolate, abaxially sericeous-villous, later glabrescent, margin often entire. |
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21 S. velutina |
| + |
Petioles 6–10 mm; leaf blades broadly ovate or rhombic-ovate, abaxially sericeous-villous, not glabrescent, margin lobed. |
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22 S. lobulata |
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| 30 (1) |
Flowers in sessile umbels, with or without leaves at base. |
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(31) |
| + |
Flowers in pedunculate umbels or umbel-like racemes, with clustered leaves at base. |
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(42) |
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| 31 (30) |
Leaf blade margin entire or with a few crenate teeth near apex; stamens equaling petals or nearly so. |
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(32) |
| + |
Leaf blade margin serrate or slightly lobed; stamens shorter than petals. |
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(36) |
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| 32 (31) |
Branchlets conspicuously angled; flowers pinkish. |
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66 S. calcicola |
| + |
Branchlets terete; flowers white. |
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(33) |
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| 33 (32) |
Leaf blades on sterile branches usually flabellate, ± as wide as long. |
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(34) |
| + |
Leaf blades on sterile branches not flabellate, longer than wide, apex acute or obtuse. |
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(35) |
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| 34 (33) |
Branchlets glabrous; leaf blades abaxially glabrous, with glaucous bloom. |
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69 S. compsophylla |
| + |
Branchlets densely pubescent, later glabrescent; leaf blades abaxially densely pubescent. |
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70 S. aquilegiifolia |
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| 35 (33) |
Branchlets glabrous or puberulent; leaf blades 15–20 × 5–7 mm; follicles glabrous. |
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67 S. hypericifolia |
| + |
Branchlets densely pubescent, later glabrescent; leaf blades 4–14 × 3–5 mm; follicles pubescent. |
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68 S. hailarensis |
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| 36 (31) |
Leaf blade margin sharply serrulate. |
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(37) |
| + |
Leaf blade margin coarsely serrate. |
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(38) |
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| 37 (36) |
Leaf blades ovate to oblong-lanceolate, abaxially pubescent; pedicels 10–24 mm, pubescent. |
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59 S. prunifolia |
| + |
Leaf blades linear-lanceolate, abaxially glabrous; pedicels 6–10 mm, glabrous. |
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60 S. thunbergii |
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| 38 (36) |
Leaf blades glabrous on both surfaces. |
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65 S. kwangsiensis |
| + |
Leaf blades pubescent on both surfaces or glabrous adaxially when old. |
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(39) |
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| 39 (38) |
Branchlets terete, pubescent or tomentose; pedicels and hypanthium abaxially glabrous. |
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(40) |
| + |
Branchlets prominently angled, pubescent or villous; pedicels and hypanthium abaxially hairy. |
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(41) |
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| 40 (39) |
Branchlets densely tomentose; leaf blade margin usually 3-lobed, with a few large, obtuse teeth above middle, base cuneate or broadly so. |
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61 S. martini |
| + |
Branchlets pubescent; leaf blade margin not lobed, with many large, crenate teeth above base, base subrounded. |
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62 S. cavaleriei |
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| 41 (39) |
Branchlets and buds appressed villous; pedicels and hypanthium abaxially villous when young; follicles glabrous or slightly pilose on adaxial suture. |
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63 S. daochengensis |
| + |
Branchlets and buds densely pubescent; pedicels, hypanthium abaxially, and follicles pubescent. |
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64 S. muliensis |
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| 42 (30) |
Buds with 2 exterior scales. |
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(43) |
| + |
Buds with several exterior scales. |
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(46) |
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| 43 (42) |
Leaf blade margin serrate; stamens longer than petals; sepals reflexed in fruit. |
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(44) |
| + |
Leaf blade margin entire or apically crenate-serrate; stamens subequaling petals; sepals erect or spreading in fruit. |
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(45) |
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| 44 (43) |
Leaf blades oblong-elliptic to oblong-ovate, margin sharply serrate above middle or near apex; petioles 2–5 mm; stamens ca. 20. |
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55 S. flexuosa |
| + |
Leaf blades broadly ovate, margin minutely serrate or doubly serrate; petioles 4–7 mm; stamens 35–50. |
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56 S. chamaedryfolia |
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| 45 (43) |
Branchlets, leaf blades, and inflorescences glabrous; follicles glabrous or slightly hairy on adaxial suture. |
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57 S. mongolica |
| + |
Branchlets, leaf blades, and inflorescences villous; follicles pubescent. |
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58 S. mollifolia |
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| 46 (42) |
Leaf blade margin entire or apically crenate-serrate. |
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(47) |
| + |
Leaf blade margin serrate or incised, sometimes lobed. |
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(55) |
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| 47 (46) |
Leaf blades abaxially hairy; follicles hairy; sepals reflexed in fruit. |
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(48) |
| + |
Leaf blades abaxially glabrous (pubescent in S. xizangensis); follicles glabrous or pilose on adaxial suture; sepals erect, spreading, or reflexed in fruit. |
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(51) |
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| 48 (47) |
Leaf blade margin with a few obtuse teeth above middle; racemes dense. |
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49 S. tarokoensis |
| + |
Leaf blade margin entire or on sterile shoots with sharp teeth apically; racemes lax. |
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(49) |
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| 49 (48) |
Branchlets glabrous or subglabrous; leaf blades abaxially glabrous or slightly pubescent in vein axils. |
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48 S. media |
| + |
Branchlets densely villous; leaf blades abaxially densely long sericeous. |
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(50) |
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| 50 (49) |
Shrubs erect, to 2 m; leaf blades ovate-elliptic or elliptic, 1.5–4.5 cm; inflorescences 15–30-flowered; stamens longer than petals. |
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46 S. sericea |
| + |
Shrubs prostrate; leaf blades broadly ovate, 0.8–1.5 cm; inflorescences 7–15-flowered; stamens slightly shorter than petals. |
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47 S. prostrata |
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| 51 (47) |
Branchlets terete, glabrous; sepals reflexed. |
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54 S. dahurica |
| + |
Branchlets angled, pubescent; sepals erect or slightly spreading. |
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(52) |
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| 52 (51) |
Leaf blades 2–4 mm wide; veins inconspicuous. |
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(53) |
| + |
Leaf blades 4–9 mm wide, ± conspicuously 3-veined at base. |
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(54) |
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| 53 (52) |
Flowers white; styles subterminal, divergent. |
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50 S. alpina |
| + |
Flowers pink-red; styles abaxial, suberect. |
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51 S. xizangensis |
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| 54 (52) |
Inflorescences 2–4 cm; styles terminal, divergent. |
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52 S. myrtilloides |
| + |
Inflorescences 1–1.5 cm; styles abaxial. |
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53 S. ningshiaensis |
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| 55 (46) |
Stamens longer than or rarely subequaling petals; flowers in umbel-like racemes. |
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(56) |
| + |
Stamens shorter than or subequaling petals; flowers in umbels. |
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(61) |
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| 56 (55) |
Follicles pubescent or hirsute; sepals erect in fruit. |
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(57) |
| + |
Follicles glabrous or subglabrous; sepals reflexed in fruit. |
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(59) |
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| 57 (56) |
Pedicels and hypanthium abaxially pubescent; leaf blades abaxially pubescent, margin lobed. |
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40 S. sublobata |
| + |
Pedicels and hypanthium abaxially glabrous; leaf blades abaxially pubescent on veins or in vein axils, margin not lobed. |
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(58) |
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| 58 (57) |
Leaf blades oblong-elliptic to lanceolate-elliptic, abaxially pubescent in vein axils, base cuneate, margin irregularly incised serrate, sometimes doubly serrate above middle. |
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41 S. elegans |
| + |
Leaf blades triangular-ovate or lanceolate-ovate, abaxially pubescent on veins, base truncate, rarely subrounded, margin doubly serrate except near base. |
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42 S. anomala |
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| 59 (56) |
Leaf blades ovate to elliptic-ovate, glabrous or abaxially with tufted hairs in vein axils, margin singly serrate, leaves on sterile shoots often incised doubly serrate. |
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43 S. laeta |
| + |
Leaf blades oblong or ovate, margin with a few coarse teeth. |
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(60) |
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| 60 (59) |
Leaf blades sericeous-villous on both surfaces, margin with 3–7 large, acute teeth on each side above middle; inflorescences 12–20-flowered; flowers 8–10 mm in diam. |
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44 S. papillosa |
| + |
Leaf blades glabrous on both surfaces, margin with 3–5 inconspicuous, triangular, obtuse teeth on each side above middle; inflorescences 5–10-flowered; flowers 10–12 mm in diam. |
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45 S. lichiangensis |
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| 61 (55) |
Leaf blades, inflorescences, and follicles glabrous. |
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(62) |
| + |
Leaf blades hairy abaxially. |
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(65) |
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| 62 (61) |
Leaf blade apex acute. |
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(63) |
| + |
Leaf blade apex obtuse. |
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(64) |
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| 63 (62) |
Leaf blade rhombic-lanceolate to rhombic-oblong, pinnately veined, margin serrate above middle. |
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36 S. cantoniensis |
| + |
Leaf blade rhombic-ovate, inconspicuously 3-veined or pinnately veined, margin usually 3-lobed, incised doubly serrate. |
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37 S. ×vanhouttei |
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| 64 (62) |
Leaf blades suborbicular, 3–5-veined, base rounded to subcordate, rarely cuneate, margin with a few crenate teeth above middle, apex usually 3-lobed. |
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38 S. trilobata |
| + |
Leaf blades rhombic-ovate to obovate, inconspicuously 3-veined or pinnately veined, base cuneate, margin with a few crenate, incised teeth or 3–5 lobes above middle. |
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39 S. blumei |
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| 65 (61) |
Inflorescences glabrous; follicles glabrous except on adaxial suture. |
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35 S. pubescens |
| + |
Inflorescences and follicles hairy. |
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(66) |
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| 66 (65) |
Leaf blades glabrous adaxially, pilose or subglabrous abaxially. |
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34 S. siccanea |
| + |
Leaf blades pilose adaxially. |
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(67) |
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| 67 (66) |
Leaf blades pubescent abaxially. |
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(68) |
| + |
Leaf blades tomentose abaxially. |
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(69) |
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| 68 (67) |
Leaf blades abaxially sericeous-pubescent, margin coarsely obtusely dentate, apex usually 3-lobed; inflorescences 2–3 cm in diam.; pedicels 6–10 mm. |
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29 S. nishimurae |
| + |
Leaf blades abaxially sparsely pubescent, margin obtusely or sharply serrate above middle or near apex, apex not 3-lobed; inflorescences 4–5 cm in diam.; pedicels 12–22 mm. |
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30 S. hirsuta |
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| 69 (67) |
Sepals ovate-lanceolate; leaf blades abaxially densely yellow tomentose, deeply sharply incised serrate. |
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31 S. chinensis |
| + |
Sepals triangular to ovate-triangular; leaf blades abaxially densely white tomentose, shallowly obtusely serrate. |
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(70) |
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| 70 (69) |
Leaf blades rhombic-ovate, 2–4.5 cm, margin obtusely incised serrate or lobed, apex acute, rarely obtuse. |
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32 S. dasyantha |
| + |
Leaf blades ovate to obovate, 1–2 cm, margin ± obtusely serrate or doubly serrate, apex obtuse or slightly 3-lobed. |
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33 S. yunnanensis |