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FOC | Family List | FOC Vol. 17 | Verbenaceae

11. Vitex Linnaeus, Sp. Pl. 2: 638. 1753.

牡荆属 mu jing shu

Trees or shrubs. Branches glabrous or sparsely pubescent. Leaves opposite, palmately (1-)3-8-foliolate; leaflets petiolulate, margin entire, dentate, serrate, or incised. Inflorescences terminal or axillary cymes, thyrses, or panicles; bracts usually small, often early deciduous. Calyx campanulate, tubular, or funnelform, sometimes 2-lipped, usually truncate or shortly 5-dentate. Corolla blue, white, or yellow, 2-lipped, lower lip 3-lobed with middle lobe greatly elongated, upper lip usually 2-lobed. Stamens 4, didynamous, sometimes exserted; anther locules attached only at tip, becoming divaricate. Ovary 2-4-locular; ovules 1 or 2 per locule. Style filiform; stigma 2-cleft. Drupes subtended by enlarged calyx, globose, ovoid, or obovoid, normally 4-locular and 4-seeded but often some locules suppressed and base of pyrene forming a hollowed cavity, endocarp a bony pyrene, mesocarp generally fleshy. Seeds obovoid or oblong, endosperm absent; cotyledons usually fleshy.

About 250 species: chiefly tropical, few in temperate regions of both hemispheres; 14 species in China.


1 Inflorescences axillary, subtended by normal leaves.   (2)
+ Inflorescences terminal, not subtended by normal leaves.   (6)
       
2 (1) Leaflets glabrous; inflorescences thyrses with a well-defined main axis and short lateral cymes   10 Vitex peduncularis
+ Leaflets pubescent; inflorescences cymes, usually rounded.   (3)
       
3 (2) Leaves often with more than 3 leaflets; cymes to 3, produced in succession from 1 axil   11 Vitex burmensis
+ Leaves 3- or 5-foliolate; cymes rarely more than 1 per axil.   (4)
       
4 (3) Corolla yellow, tube slender and cylindric   12 Vitex vestita
+ Corolla white flushed with pink or blue, tube funnelform to campanulate.   (5)
       
5 (4) Cyme branches narrowly divergent; corolla to 1.7 cm   13 Vitex yunnanensis
+ Cyme branches widely divergent; corolla to 1 cm   14 Vitex duclouxii
       
6 (1) Peduncles, pedicels, and calyces glabrous or subglabrous.   (7)
+ Peduncles, pedicels, and calyces pubescent.   (9)
       
7 (6) Leaflets serrate; calyx distinctly dentate   3 Vitex sampsonii
+ Leaflets entire; calyx teeth rudimentary.   (8)
       
8 (7) Branchlet lenticels obscure; corolla outside puberulent   1 Vitex kwangsiensis
+ Branchlet lenticels conspicuous; corolla outside glabrescent   2 Vitex tripinnata
       
9 (6) Leaflets abaxially pubescent only along main veins.   (10)
+ Leaflets abaxially densely pubescent.   (11)
       
10 (9) Leaflets lanceolate to oblong-lanceolate; bractlets persistent in flower   4 Vitex pierreana
+ Leaflets obovate to elliptic or oblong; bractlets early deciduous   5 Vitex quinata
       
11 (9) Leaflets with glandular hairs, adaxially glabrescent and grayish dotted; bracts deciduous   6 Vitex canescens
+ Leaflets puberulent, not adaxially dotted; bracts persistent.   (12)
       
12 (11) Prostrate shrubs   8 Vitex rotundifolia
+ Erect shrub or small trees.   (13)
       
13 (12) Leaves (1-)3(-5)-foliolate; middle leaflet sessile; calyx with 5 obscure teeth, shorter than fruit   7 Vitex trifolia
+ Leaves 3-7-foliolate; middle leaflet distinctly petiolulate; calyx with 5 distinct teeth, as long as fruit   9 Vitex negundo

  • List of lower taxa


     

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