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26. Yinshania Ma & Y. Z. Zhao, Acta Phytotax. Sin. 17(3): 113. 1979.
阴山荠属 yin shan ji shu
Cochleariella Y. H. Zhang & R. Vogt; Cochleariopsis Y. H. Zhang (1985), not Á.. Löve & D. Löve (1976); Hilliella (O. E. Schulz) Y. H. Zhang & H. W. Li.
Herbs annual or perennial with tubers or rhizomes. Trichomes absent or simple or 1- or 2-forked. Stems erect, ascending, or decumbent. Basal leaves not rosulate, simple, undivided or pinnately or palmately lobed, trifoliolate, or pinnately compound. Cauline leaves petiolate, not auriculate, similar to basal leaves or simple upward. Racemes few to many flowered, ebracteate or rarely bracteate throughout. Fruiting pedicels erect, ascending, divaricate, or reflexed. Sepals oblong or ovate, spreading, not saccate. Petals white or rarely pinkish, spreading, longer than sepals; blade obovate, oblong, or ovate, apex rounded; claw absent or short. Stamens 6, spreading, subequal; filaments not dilated at base; anthers ovate or oblong, apex obtuse. Median nectaries absent; lateral glands 1 on each side of lateral stamen. Ovules 1-24 per ovary. Fruit dehiscent silicles, oblong, ovoid, ellipsoid, or globose, terete, rarely slightly latiseptate or angustiseptate, sessile; valves veinless or obscurely veined, glabrous, papillate, or with simple trichomes, smooth; replum rounded; septum complete, perforated, or absent; style obsolete or to 3 mm; stigma discoid, entire. Seeds uniseriate or biseriate, wingless, oblong, ovate, reniform, orbicular, ovoid, lenticular, elliptic, ellipsoid, or globose, slightly flattened; seed coat minutely reticulate or papillate, not mucilaginous when wetted; cotyledons incumbent or rarely accumbent.
Thirteen species: China, N Vietnam; 13 species (12 endemic) in China.
All species of Yinshania were treated in FRPS as members of Cochlearia, a genus that does not occur in China, the Himalaya, or SW Asia. A few species of Cochlearia grow in N Russia, with the range of one extending into Japan. The record of C. officinalis Linnaeus in FPRS (p. 98) is likely an error. For a detailed account of Yinshania and its segregates see Al-Shehbaz et al. (Harvard Pap. Bot. 3(1): 79-94. 1998).
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| 1 |
Most or all leaves simple, entire, repand, or sinuate, rarely lowermost few trifoliolate |
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8 Y. sinuata |
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Leaves compound or pinnately lobed. |
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(2) |
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| 2 (1) |
Flowers bracteate along entire length or only on lower half of inflorescence. |
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(3) |
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Flowers ebracteate. |
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(4) |
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| 3 (2) |
Plants annual; ovary ca. 8-ovuled, cylindric, papillate |
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13 Y. hui |
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Plants perennial; ovary ca. 4-ovuled, obovate-ellipsoid, glabrous |
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12 Y. hunanensis |
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| 4 (2) |
Trichomes forked. |
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(5) |
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Trichomes simple or absent. |
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(6) |
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| 5 (4) |
Leaves and stems canescent, densely villous with trichomes 0.3-0.5(-0.6) mm |
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6 Y. furcatopilosa |
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Leaves and stems not canescent, puberulent with trichomes 0.05-0.08(-0.1) mm |
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3 Y. zayuensis |
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| 6 (4) |
Infructescence rachis conspicuously flexuous. |
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(7) |
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Infructescence rachis straight. |
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(8) |
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| 7 (6) |
Leaves canescent, densely villous, margin of lower leaves deeply crenate; seeds 0.7-0.9 mm, (4-)10-20 per fruit |
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5 Y. henryi |
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Leaves not canescent, glabrous or sparsely pubescent, margin of lower leaves not deeply crenate; seeds 1.5-2 mm, 2(-4) per fruit |
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2 Y. yixianensis |
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| 8 (6) |
Leaflets of middle and lower leaves less than 1 cm wide. |
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(9) |
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Leaflets of middle and lower leaves more than 1 cm wide. |
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(10) |
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| 9 (8) |
Leaves glabrous, lower ones trifoliolate; fruit papillate |
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4 Y. fumarioides |
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Leaves usually pilose, rarely glabrous, lower ones pinnate, with 2-5 pairs of lateral leaflets; fruit glabrous or rarely pilose |
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1 Y. acutangula |
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| 10 (8) |
Lower leaves with 5-9 leaflets; fruit conspicuously compressed; plants tuberous perennials |
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11 Y. rupicola |
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Lower leaves with 3(or 5) leaflets; fruit plump, not compressed; plants annuals with slender roots. |
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(11) |
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| 11 (10) |
Leaflet apex acuminate-caudate; fruit appressed to rachis; plants puberulent with trichomes to 0.1 mm |
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10 Y. lichuanensis |
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Leaflet apex obtuse or acute, rarely subacuminate; fruit widely spreading or reflexed; plants glabrous throughout or rarely glabrescent. |
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(12) |
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| 12 (11) |
Fruit ovate or obovate, 2- or 3-seeded; style 0.5-0.8 mm; pedicel (1.5-)2.5-4 mm |
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9 Y. paradoxa |
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Fruit narrowly oblong, 7-10-seeded; style (1-)1.5-2 mm; pedicel 6-10 mm |
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7 Y. rivulorum |
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List of lower taxa
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