细齿角蕨 xi chi jiao jue

Phegopteris crenulatoserrulata Makino, Bot. Mag. (Tokyo) 17: 78. 1903; Athyrium austro-ussuriense (Komarov) Fomin; A. crenulatoserrulatum Makino; A. koryoense C. Christensen; Cornopteris coreana Nakai; C. crenulatoserrulata f. coreana (Nakai) H. Itô; Dryopteris austro-ussuriensis Komarov; D. crenulatoserrulata (Makino) C. Christensen; Neoathyrium crenulatoserrulatum (Makino) Ching & Z. R. Wang; Phegopteris austro-ussuriensis (Komarov) Komarov & Alissova; Pseudathyrium crenulatoserrulatum (Makino) Nakai.

Plants terrestrial, medium-sized. Rhizome moderately stout, creeping, apex sparsely scaly; scales brownish, ovate-lanceolate, membranous, large, up to 1 cm or longer. Fronds distant; stipe nearly as long as or slightly longer than lamina, 40-60 cm, basal part moderately stout with pneumatophores on lateral ridges, base not beak-shaped, up to 7-9 mm in diam., scaly like rhizome apex; lamina 3-pinnatifid, deltoid-ovate to ovate-oblong, 25-70 × 20-60 cm, base broadly cuneate or somewhat cordate, apex acuminate; pinnae 10-15 pairs, broadly lanceolate or oblong-lanceolate, lower pinnae with stalks 2-10 mm, subopposite, ascending; basal 2 pairs of pinnae largest, 10-32 × 4-8 cm, base slightly narrowed, broadly cuneate or subtruncate, apex acuminate; upper pinnae gradually smaller, lanceolate, alternate, subsessile or sessile; pinnules 8-20 pairs, catadromous, lanceolate or oblong-lanceolate, 1-4 cm × 5-15 mm, base broadly cuneate, subsessile, apex acuminate; lower pinnules subopposite; upper pinnules alternate, spreading or slightly ascending; ultimate segments 5-10 pairs, oblong, base adnate to narrow costular wing, crenulate at margin, apex obtuse-rounded. Veins inconspicuous adaxially, visible abaxi-ally, pinnate, midrib slightly tortuous, veinlets simple or forked. Lamina papery when dried, greenish brown adaxially, green abaxially; fleshy slender hornlike processes with scaly apices present in adaxial groove of costules at bases of costules; rachis, costae, and costules abaxially with pallid unicellular short hairs and pale brown multicellular long hairs. Sori orbicular or elliptic, exindusiate, abaxial, medial. Spores bilateral, perispore with rugate processes. 2n = 80*.

Subalpine mixed forests, grasslands; 800-1000 m. Heilongjiang, Henan, Jilin, Liaoning, Shaanxi [Japan, Korea, E Russia].