直立对囊蕨 zhi li dui nang jue

Athyriopsis erecta Z. R. Wang, Acta Phytotax. Sin. 20: 237. 1982; Dryoathyrium articulatipilosum Ching & W. M. Chu ex Y. T. Hsieh; D. chingii (Z. Y. Liu) W. M. Chu ex P. S. Wang & X. Y. Wang; D. erectum (Z. R. Wang) W. M. Chu & Z. R. Wang; Lunathyrium chingii Z. Y. Liu.

Rhizomes creeping, ascending at apex. Fronds caespitose; fertile fronds up to 55-113 cm; stipe upward brownish stramineous, 30-40(-68) cm, 2-3(-4.2) mm in diam., base not beak-shaped, with sparse brown, ovate or broadly lanceolate scales, upper part glabrate; lamina 2-pinnatipartite, narrowly ovate or deltoid-ovate, 25-40(-48) × 14-24 cm, base rounded-cuneate, apex acuminate; pinnae 5-8(-12) pairs, lower 1-6 pairs subopposite, basal 1 or 2 pairs nearly equally long, shortly stalked, stalks up to 1.5 cm; upper pinnae alternate, ascending or subspreading, 2-5 cm apart, falcate-lanceolate; middle and lower pinnae 10-14.5 × 2.5-4 cm, subequilateral, base truncate, apex acuminate; basal pinnae prominently narrowed and slightly inequilateral to base, pinnatipartite; segments 10-15(-17) pairs, basal segments subopposite, others alternate, slightly ascending, approximate, oblong or rectangular, 1-2 cm × 6-11 mm, shallowly toothed on lateral margin, obtuse-rounded or subtruncate at apex. Veins pinnate in segment, lateral veins 6-9 pairs, simple or forked, occasionally 3- or 4-forked. Lamina herbaceous when dried, greenish brown; rachis and costae with sparse brown lanceolate small scales and scalelike hairs. Sori oblong, occasionally shortly linear, J-shaped, or horseshoe-shaped, 1-3 mm, abaxial, medial, 1-7 pairs per segment; indusia brownish, oblong, occasionally reniform or horseshoe-shaped, basal acroscopic one often hooked, membranous, erose at margin, persistent. Perispore with long spines. 2n = 80*.

● Valley forests, wet areas; (200-)1200-2500 m. Chongqing, Guizhou, Hubei, Hunan, Sichuan, NE Yunnan.

The long spines on the perispore of Deparia erecta are unique in the genus.