柳羽鳞毛蕨 liu yu lin mao jue

Dryopteris submarginata Loyal (1965), not Rosenstock (1914).

Plants more than 1 m tall. Rhizome creeping. Stipe stramineous, ca. 1/2 as long as lamina, base densely clothed with ovate, pale brown, adnate scales; lamina deltoid-lanceolate, ca. 70 cm, widest at base, 40-50 cm wide, twice pinnate-pinnatifid; pinnae ca. 20 pairs, oblong-lanceolate, basal pair ca. 25 cm, ca. 12 cm wide at base, bipinnate, becoming tripinnatifid deeply or halfway, apex acuminate, stalk ca. 1 cm, pinnae sessile upward; pinnules ca. 12 pairs, oblique, narrowly oblong-lanceolate, basal pinnule ca. 6 cm, 1-2 cm wide at base, cuneate, shortly stalked, apex acuminate, other pinnules sessile, pinnatifid (pinnatipartite at base); segments (ultimate pinnules) 6-8 pairs, oblique, deltoid-oblong, margin serrate, generally revolute, apex obtuse or truncate, sharply deltoid-serrate. Lamina thinly leathery; rachis and costa glabrous or sparsely scaly; scales pale brown, lanceolate and nodose, moniliform; veins pinnate, impressed adaxially, raised abaxially. Upper part of lamina fertile, sori in 1 regular row on each side of costule and close to costule; indusia ferruginous, orbicular-reniform, thick, persistent; spores ellipsoid, with wrinkled epispore, forming slices or papilla protuberance.

Forests; 2000-2400 m. W Yunnan [Bhutan, India, Nepal].