崖县叶下珠 ya xian ye xia zhu

Phyllanthus annamensis Beille; P. campanulatus Ridley; P. coriaceus Wallich ex J. D. Hooker; P. frondosus Wallich ex Müller Argoviensis var. rigidus Ridley; P. klossii Ridley; P. sciadiostylus Airy Shaw.

Erect shrubs up to 4 m tall, glabrous throughout; stem gray-brown; branchlets sharply 4-angled. Stipules ovate-triangular, ca. 1 mm; petiole 2-3 mm; leaf blade ovate-lanceolate or rhombic, 3-14 × 1.5-3.5 cm, thinly leathery, base broadly cuneate, margin revolute when dry, apex acuminate, slightly caudate; lateral veins (4-)6-8 pairs. Plants monoecious. Inflorescence a bisexual fascicle, axillary or along specialized leafless branchlets, 2-10 cm, usually with 1-3 female and 2-6 male flowers, with many 2-3 mm scaly bracts. Male flowers ca. 4 mm in diam.; pedicels 1-2(-3) mm; sepals 6, obovate, subequal, ca. 1.5 mm, apex obtuse or acute, margin membranous; stamens 3; filaments connate, ca. 1 mm; thecae parallel, longitudinally dehiscent; pollen grain spherical or oblate-spherical, 6-colporate, rarely 4- or 5-colporate or forate-colpate; disk glands 6. Female flowers ca. 4 mm in diam.; pedicels 1-3.5 mm; sepals 6, obovate, ca. 1.5 × 1.5 mm, margin membranous, persistent in fruit; disk cup-shaped, membranous, slightly lacerate at apex; ovary ovoid, ca. 1.3 mm in diam., 3-celled; styles 3, ca. 1 mm, connate at base, bifid at apex, lobes recurved and slightly appressed to apex of ovary. Fruiting pedicel ca. 4 mm; capsule globose, ca. 4 mm in diam., red. Seed red, ca. 2 mm in diam. Fl. Mar-Jun, fr. May-Sep.

Open montane forests, disturbed scrub, riversides; 100-600 m. S Hainan (Sanya) [Malaysia, Thailand, Vietnam].