140. Dryopteris ×holttumii Li Bing Zhang
霍氏鳞毛蕨 huo shi lin mao jue
Replaced synonym: Dryopsis ×fauriei Holttum & P. J. Edwards, Kew Bull. 41: 198. 1986; Dryopteris ×fauriei (Holttum & P. J. Edwards) Li Bing Zhang, Phytotaxa 71: 21. 2012, not D. fauriei Kodama, Icon. Pl. Koisikav. 2: 11. 1914.
Rhizomes erect. Stipe castaneous, > 30 cm; stipe scales dark brown, lanceolate, ca. 8 × 1.5 mm. Lamina 2-pinnate to 3-pinnatifid, lanceolate, ca. 90 × 50 cm; free lateral pinnae ca. 20 pairs, continuous; basal pinnae not shortened, ca. 26 × 4.5 cm, not reflexed; middle pinnae ca. 26 × 4.5 cm; pinnules entire to deeply lobed, hyaline margins lacking, obtuse at apex. Rachises and costae abaxially scaly, scales on costae ovate, costules without scales. Veins of ultimate pinnules pinnate. Sori confined to distal parts of pinnules. Indusia distinct and persistent. Spores ?not developed.
● Broad-leaved and mixed forests; 2000-2800 m. Taiwan (Jiayi, Nantou, Yilan).
Dryopteris ×holttumii is known from three counties in Taiwan. It is presumably a hybrid between D. apiciflora and D. maximowicziana (Holttum & Edwards, loc. cit.: 199, as Dryopsis maximowicziana). The latter hypothetical parent was originally not found in Taiwan but is now rediscovered there, although, according to reviewer Ralf Knapp, Dryopteris ×holttumii and D. maximowicziana have not been observed to occur at the same locations together.