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FOC | Family List | FOC Vol. 2-3 | Dryopteridaceae | Dryopteris

160. Dryopteris hookeriana (T. Moore) Li Bing Zhang, Taxon. 61: 1210. 2012.

虎克鳞毛蕨 hu ke lin mao jue

Diacalpe hookeriana T. Moore, Gard. Chron. 1854: 135. 1854; D. aspidioides Blume var. hookeriana (T. Moore) Ching & S. K. Wu.

Rhizome short, erect, ca. 1 cm in diam., terminal part densely covered with dark brown scales; scales ovate-lanceolate, 8-10 mm, thickly membranous, entire, acuminate. Fronds tufted; stipe 25-32 cm, base 2.5-3.5 mm in diam., dark brown, base covered with scales same as on rhizome, smaller and sparser upward; lamina dark brown when dry, ovate or narrowly ovate, 30-50 × 16-40 cm, quadripinnatifid, papery, with darkish red and globose glands along veins abaxially, sparsely covered with short nodose hairs adaxially, apex long acuminate; pinnae 16-20 pairs, basal ones subopposite, 5-7 cm apart, subsessile, upper ones alternate, close together, basal pairs larger, triangular-lanceolate, falcate, 15-25 × 6.5-11 cm, tripinnatifid, base rounded-truncate, apex caudate-acuminate; pinnules 16-18 pairs, always alternate, anadromous, subsessile, 1.3-2.5 cm apart, basiscopic pinnule longest, lanceolate, 5-10 × 1.5-2.6 cm, bipinnatifid, base symmetrical, rounded-truncate; ultimate pinnules 10-17 pairs, subsessile, separate, elliptic, 1-2 cm, base asymmetrical, broadly cuneate, pinnatipartite to ultimate rachillae, apex with 2-4 small teeth; lobes 4 or 5 pairs, basal pair opposite, distant, elliptic, 3-6 × 1-2.2 mm, base asymmetrical, apex rounded, with 2-4 small teeth or entire; veins obvious abaxially, pinnate, veinlets simple or furcate, submarginal; rachis and rachillae dark brown, sparsely covered with brown scales and curly nodose hairs. Sori globose, ca. 0.8 mm in diam., 1 in each ultimate pinnule or lobe, dorsal on basal acroscopic veinlets; indusium inferior, brown, leathery, dehisced into 2 or 3 irregular valves from top when mature; spores narrowly elliptic, with fold.

Broad-leaved forests; 1600-2200 m. Xizang [India, Nepal].

None of the following related names are validly published: "Diacalpe aspidioides var. himalayensis" (Ching & S. K. Wu, Acta Phytotax. Sin. 21: 376. 1983, nom. nud.), "Diacalpe aspidioides var. hookeriana Wallich ex Ching & S. H. Wu" (Acta Phytotax. Sin. 21: 376. 1983, nom. nud.), and "Sphaeropteris hookeriana" (Wallich, Numer. List, no. 248. 1829, nom. nud.).

This species was sometimes treated as a variety of Dryopteris pseudocaenopteris. Dryopteris hookeriana differs from the latter in its subsessile pinnae and pinnules.


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