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FOC | Family List | FOC Vol. 2-3 | Dryopteridaceae | Dryopteris

165. Dryopteris annamensis (Tagawa) Li Bing Zhang, Taxon. 61: 1210. 2012.

中越鳞毛蕨 zhong yue lin mao jue

Diacalpe annamensis Tagawa, Acta Phytotax. Geobot. 14: 46. 1950; D. caudifolia Ching & S. K. Wu.

Rhizome short, erect, woody, 1-1.2 cm in diam., apex densely covered with dark brown scales; scales lanceolate, 0.6-0.8 cm, entire, acuminate. Fronds tufted; stipe 24-36 cm, base 2.5-3.5 mm in diam., dark brown, densely covered with scales as on rhizome, sparser and smaller upward; lamina brown or green when dry, ovate, 25-35 × 20-27 cm, quadripinnatifid, papery, sometimes with citrus-colored glands along veins abaxially, sparsely covered with brown, short nodose hairs adaxially, apex long acuminate; pinnae 19-22 pairs, subopposite, basal ones with short stalk 0.3-1.1 cm, 4.5-6 cm apart from second pair, upper ones subsessile, basal pair largest, triangular-lanceolate, 14-16 × 8-10 cm, tripinnatifid, base nearly rounded-truncate, apex acuminate; pinnules 16-22 pairs, alternate, anadromous, with short stalk 0.5-2 mm, 1-2 cm apart, close together, basiscopic 1 or 2 pinnules usually enlarged, lanceolate, 6.5-7 × 2.2-2.7 cm, bipinnatifid, base asymmetrical, apex acute and undulate; ultimate pinnules 14-16 pairs, basal 1 or 2 pinnules opposite, alternate upward, with short stalk ca. 1.2 mm, distant, elliptic, 1-1.5 × 0.4-0.8 cm, base asymmetrical, pinnatipartite to ultimate rachis, apex rounded; lobes 3-5 pairs, distant, elliptic, 3.5-5 × 1.7-2 mm, entire; median pinnae narrower than other pinnae, basiscopic pinnules longer than upper ones, catadromous, ultimate pinnules entire or dentate-lobed; rachis and rachillae stramineous, sparsely covered with brown nodose hairs and small scales. Veins obvious abaxially, brown, pinnate on lobes, veinlets simple, submarginal. Sori globose, ca. 1 mm in diam., enveloped by indusium, 1 in each ultimate pinnule or lobe, dorsal on basal acroscopic veinlets; indusium brown, globose, leathery, dehisced into 2 or 3 irregular valves from top when mature; spores narrowly elliptic.

Dense forests; 1600-2800 m. Hainan, Taiwan, Xizang, Yunnan [Vietnam].


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