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19. CAREX sect. Mitratae Kükenthal in Engler, Pflanzenr. 38(IV. 20): 458. 1909.

灰帽薹草组 hui mao tai cao zu

Authors: Yancheng Tang, Shuren Zhang, Tetsuo Koyama & Gordon C. Tucker

Spikes usually unisexual; terminal spike male; lateral spikes often female, rarely androgynous, female spike long cylindric, cylindric, or linear. Utricles herbaceous, glabrous or puberulent, shortly beaked or nearly beakless, orifice truncate or 2-toothed. Nutlet tightly enveloped, trigonous, sometimes with contracted angles and excavated faces, apex often discoid-annulate; stigmas 3.

About 60 species: C, E, and SE Asia, Australia, Europe, Pacific islands (New Zealand); 27 species (16 endemic) in China.


1 Nutlet contracted on angles or excavated at faces   (2)
+ Nutlet neither contracted on angles nor excavated at faces   (11)
       
2 (1) Nutlet contracted on angles   (3)
+ Nutlet not contracted on angles   (6)
       
3 (2) Nutlet excavated at faces.   103 C. chungii
+ Nutlet not excavated at faces   (4)
       
4 (3) Leaves not trabeculate; utricles 2-2.5 mm.   94 C. foraminata
+ Leaves trabeculate; utricles 3.5-5 mm   (5)
       
5 (4) Peduncles of female spikes mostly included; nutlets ca. 2 mm.   97 C. tungfangensis
+ Peduncles of female spikes mostly exserted; nutlets ca. 4 mm.   95 C. tianmushanica
       
6 (2) Utricle distinctly convex adaxially, beak recurved; nutlet only slightly excavated at lower part of face.   98 C. foraminatiformis
+ Utricle indistinctly convex adaxially, beak erect; nutlet excavated at both upper and lower part of face   (7)
       
7 (6) Leaf sheaths and blades puberulent.   102 C. duvaliana
+ Leaf sheaths and blades glabrous   (8)
       
8 (7) Utricles ovate-fusiform, 2.5-4 mm.   100 C. dolichostachya
+ Utricles elliptic-rhombic, oblong-ellipsoid, or rhombic-oblong, 2.5-3 mm   (9)
       
9 (8) Male spike 25-45 mm; nutlet 2.5-3 mm.   96 C. qingliangensis
+ Male spike 10-30 mm; nutlet 1.5-2 mm   (10)
       
10 (9) Female spike linear-cylindric, 2-2.5 mm wide, loosely flowered.   101 C. rugata
+ Female spike cylindric or oblong-cylindric, 3-4 mm wide, densely flowered.   99 C. sociata
       
11 (1) Utricles lanceolate-ovate, 3.5-4.5 mm   (12)
+ Utricles ellipsoid, ovoid, or ovate-fusiform, 2-3(-4) mm   (13)
       
12 (11) Spikes usually 2; terminal male spike ovate-clavate; lateral spike androgynous.   107 C. shanghangensis
+ Spikes 2-4; terminal male spike linear-oblong; lateral spikes female.   108 C. orthostemon
       
13 (11) Lower involucral bracts hardly or shortly sheathing (3-7 mm)   (14)
+ Lower involucral bracts long sheathing (1-3 cm)   (19)
       
14 (13) Male spike shortly pedunculate, base not surpassing next female spike   (15)
+ Male spike long pedunculate, base surpassing next female spike   (16)
       
15 (14) Male spike pale brown; female spikes linear to linear-cylindric, 2-3 mm wide.   106 C. mitrata
+ Male spike pale green; female spikes ellipsoid to ovoid, more than 3 mm wide.   109 C. breviculmis
       
16 (14) Utricles rhombic-elliptic to broadly ovate or obovate   (17)
+ Utricles ovate-elliptic or ovate-oblong   (18)
       
17 (16) Utricles puberulent, stipe very short and not conspicuous.   110 C. hypochlora
+ Utricles only sparsely puberulent, stipe conspicuous though short.   113 C. subebracteata
       
18 (16) Rhizome slender, long creeping; costa of female glumes not excurrent beyond apex.   119 C. nervata
+ Rhizome short; plants densely tufted; costa of female glumes excurrent into a mucro.   120 C. kiangsuensis
       
19 (13) Male glumes usually connate at margin forming a cuplike structure; most spikes fastigiate to subfastigiate; male spike linear   (20)
+ Male glumes scalelike, not connate at margin; spikes racemose; male spike ovate-cylindric   (21)
       
20 (19) Leaves slightly verruculose on upper surface; male glumes with margins connate.   105 C. pseudotristachya
+ Leaves not verruculose on upper surface; male glumes with margins connate or free.   104 C. tristachya
       
21 (19) Female spike ovate, ovate-oblong, or cylindric, 4-5 mm thick, with denser utricles   (22)
+ Female spike linear to linear-cylindric, rarely oblong-ovate, 2-3 mm thick, with rather lax utricles   (24)
       
22 (21) Utricle nearly beakless, orifice entire.   111 C. davidii
+ Utricle shortly beaked, orifice minutely or acutely 2-toothed   (23)
       
23 (22) Female glumes pale; utricle sparsely puberulent.   112 C. breviaristata
+ Female glumes pale or stramineous and pale brown on upper margins; utricle glabrous.   114 C. qiyunensis
       
24 (21) Leaf blade hairlike, up to 2 mm wide.   115 C. capilliformis
+ Leaf blade 2-4 mm wide   (25)
       
25 (24) Beak of utricle ± recurved, orifice toothless.   116 C. polyschoenoides
+ Beak erect, orifice 2-toothed   (26)
       
26 (25) Utricle densely puberulent; female glumes with mucro; nutlet gray-black, ovate, shortly stipitate.   117 C. pisiformis
+ Utricle glabrous; female glumes without mucro; nutlet ashy brown, ellipsoid, not stipitate.   118 C. hezhouensis

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