1 |
Radical leaves pinnate (sometimes 3-foliolate in P. hypargyrea). |
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(2) |
+ |
Radical leaves palmately 3–5-foliolate. |
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(24) |
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2 (1) |
Leaves abaxially green or greenish, sericeous, pilose, or glabrescent. |
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(3) |
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Leaves abaxially densely white or yellowish tomentose or sericeous. |
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(9) |
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3 (2) |
Leaflet margin parted to midvein. |
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(4) |
+ |
Leaflet margin serrate or lobed, but not deeply so to midvein. |
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(6) |
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4 (3) |
Radical leaves 3-foliolate, or terminal leaflets pseudopinnately parted, abaxially white sericeous. |
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62 P. hypargyrea |
+ |
Radical leaves pinnate, abaxially green, pilose or sparsely sericeous. |
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(5) |
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5 (4) |
Leaflets 2-pinnately dissected, ultimate segments ca. 1 mm wide, both surfaces pilose, apex acuminate; stipules proximally adnate to petiole for ca. 3/4 their length; inflorescences 1–3-flowered. |
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61 P. coriandrifolia |
+ |
Leaflets 1-pinnatifid, ultimate segments ca. 2 mm wide, both surfaces pilose and granular sessile glandular, apex obtuse; stipules proximally adnate to petiole for ca. 1/2 their length; inflorescences cymose, many flowered. |
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63 P. granulosa |
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6 (3) |
Radical leaves with 2 or 3(or 4) pairs of leaflets. |
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(7) |
+ |
Radical leaves with 4–23 pairs of leaflets. |
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(8) |
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7 (6) |
Leaflet margin coarsely acutely or obtusely incised serrate; flowers ca. 2 cm in diam.; epicalyx segments equaling sepals. |
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66 P. lancinata |
+ |
Leaflet margin crenulate; flowers 1.5–1.8 cm in diam.; epicalyx segments shorter than sepals. |
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67 P. crenulata |
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8 (6) |
Inflorescence lax, many flowered; flowers usually 1–1.5 cm in diam.; epicalyx segments linear- lanceolate, usually shorter than sepals, not enlarged in fruit. |
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64 P. tanacetifolia |
+ |
Inflorescence compact, few flowered; flowers 1.5–1.8 cm in diam.; epicalyx segments oblong- lanceolate, nearly equaling or slightly longer than sepals, enlarged in fruit. |
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65 P. longifolia |
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9 (2) |
Radical leaves with 2–4 pairs of leaflets, leaflet margin serrate. |
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(10) |
+ |
Radical leaves with (2 or)3–11 pairs of leaflets, leaflet margin parted or lobed. |
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(13) |
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10 (9) |
Flowering stems and petioles pilose or villous. |
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47 P. griffithii |
+ |
Flowering stems and petioles ± interwoven white tomentose and pilose, rarely glabrescent. |
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(11) |
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11 (10) |
Leaflets abaxially appressed sericeous on veins; sepals abaxially densely villous or sericeous. |
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46 P. potaninii |
+ |
Leaflets abaxially white tomentose on veins; sepals abaxially densely white tomentose. |
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(12) |
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12 (11) |
Leaflets in 2–4 pairs, oblong or oblong-lanceolate, margin incised serrate. |
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45 P. discolor |
+ |
Leaflets in 2 pairs, oblong to obovate-oblong, margin pectinately lobed. |
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48 P. hololeuca |
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13 (9) |
Flowering stems and petioles ± interwoven white tomentose or downy, rarely glabrescent. |
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(14) |
+ |
Flowering stems and petioles sericeous, villous, floccose, or pubescent, but not tomentose. |
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(17) |
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14 (13) |
Leaflets of radical leaves pinnately or palmately parted nearly to base, thereby appearing verticillate. |
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40 P. verticillaris |
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Leaflets of radical leaves not as above, not appearing verticillate. |
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(15) |
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15 (14) |
Leaflet margin pectinate, lobes fasciated oblong or oblong-lanceolate. |
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44 P. strigosa |
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Leaflet margin not pectinate, lobes often broader. |
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(16) |
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16 (15) |
Leaflets in 3–5(–8) pairs, subleathery, abaxially densely white tomentose, margin oblong or triangular lobed. |
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39 P. sischanensis |
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Leaflets in 4–8 pairs, thinly papery, abaxially canescent tomentose, margin oblong dentate or triangular lobed. |
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41 P. limprichtii |
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17 (13) |
Leaflets abaxially wholly tomentose or sericeous, sometimes glabrescent. |
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(18) |
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Leaflets abaxially tomentose, white sericeous on veins. |
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(19) |
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18 (17) |
Leaflets abaxially densely appressed sericeous, sometimes glabrescent, margin flat or slightly revolute; stipules of cauline leaves entire; flowering stems and petioles sparsely sericeous or glabrescent. |
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36 P. multiceps |
+ |
Leaflets abaxially tomentose or sericeous, margin strongly revolute; stipules of cauline leaves entire or 2-fid; flowering stems and petioles densely spreading white sericeous. |
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38 P. sericea |
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19 (17) |
Leaflets abaxially tomentose and sparsely white villous, adaxially greenish, appressed white villous or sericeous, lobes adjacent. |
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(20) |
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Leaflets abaxially sericeous only on veins, adaxially green, appressed pilose, rarely glabrescent, lobes distant. |
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(21) |
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20 (19) |
Radical leaves with 3–5 pairs of leaflets; stipules of cauline leaves entire, rarely 2-fid; stems and petioles appressed white villous. |
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35 P. pamiroalaica |
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Radical leaves with 6–9 pairs of leaflets; stipules of cauline leaves 2- or 3-fid; stems and petioles spreading white sericeous-villous. |
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37 P. plumosa |
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21 (19) |
Flowers usually 1.2–1.5 cm in diam.; sepals erect, enlarged after flowering. |
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(22) |
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Flowers usually 0.8–1.2 cm in diam.; sepals adnate to fruit, not enlarged after flowering. |
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(23) |
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22 (21) |
Leaflets usually parted almost to midvein, lobes narrow, ± fasciated lanceolate; plants appressed pubescent or recumbent sericeous, rarely glabrescent, hairs to 2 mm. |
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33 P. multifida |
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Leaflets usually lobed, but not nearly midvein, lobes triangular-oblong or triangular-lanceolate to fasciated oblong; plants sparsely spreading white villous, hairs 3–4 mm. |
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43 P. conferta |
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23 (21) |
Stipules of cauline leaves usually dentately lobed; flowering stems white sericeous. |
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42 P. chinensis |
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Stipules of cauline leaves usually entire; flowering stems white villous. |
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34 P. multicaulis |
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24 (1) |
Leaves abaxially white tomentose or sericeous. |
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(25) |
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Leaves abaxially green, pilose. |
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(37) |
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25 (24) |
Radical leaves palmately 5-foliolate, rarely 3-foliolate with leaflets pinnately parted, cauline leaves 3 to many; inflorescence corymbose, many flowered. |
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(26) |
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Radical leaves 3(or 5)-foliolate, cauline leaves very few, 1- or 2-foliolate; inflorescence 2- or 3-flowered. |
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(29) |
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26 (25) |
Leaflets of radical leaves 5–8-lobed; flowers 0.8–1 cm in diam. |
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57 P. virgata |
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Leaflets of radical leaves serrate or 1–4-lobed; flowers 1–2 cm in diam. |
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(27) |
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27 (26) |
Leaflets abaxially gray pilose or tomentose, margin flat, incised serrate. |
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60 P. inclinata |
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Leaflets abaxially densely white tomentose, margin slightly or markedly revolute, irregularly 2–5-dentate distally or 2–4-lobed on each side. |
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(28) |
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28 (27) |
Leaflet margin slightly revolute, 2–4-lobed on each side, lobes narrowly fasciated. |
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58 P. angustiloba |
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Leaflet margin markedly revolute, entire in proximal 1/2, irregularly 2–5-dentate in distal 1/2. |
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59 P. argentea |
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29 (25) |
Radical leaves 5-foliolate. |
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51 P. saundersiana |
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Radical leaves 3-foliolate. |
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(30) |
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30 (29) |
Flowering stems and petioles usually white tomentose; leaflets abaxially tomentose on lateral veins. |
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(31) |
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Flowering stems and petioles villous or also white tomentose; leaflets abaxially sericeous or pilose. |
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(32) |
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31 (30) |
Leaflets oblong-lanceolate, margin coarsely triangular 5–7-serrate; sepals abaxially appressed pilose. |
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49 P. nivea |
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Leaflets elliptic-ovate or obovate, margin sharply ovate 3–7-serrate; sepals abaxially white tomentose. |
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50 P. betonicifolia |
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32 (30) |
Stipules of cauline leaves broad, fasciated 2–4-lobed; flowers 2–3 cm in diam.; epicalyx segments abaxially silvery tomentose. |
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53 P. delavayi |
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Stipules of cauline leaves small, entire, rarely 2-serrate; flowers 1–2 cm in diam.; epicalyx segments abaxially usually green and pilose. |
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(33) |
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33 (32) |
Pedicels and sepals abaxially glandular hairy; leaflets abaxially canescent tomentose, glabrescent when old, adaxially pilose and glandular hairy. |
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55 P. evestita |
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Pedicels and sepals abaxially not glandular hairy; leaflets abaxially densely persistently tomentose, adaxially appressed sericeous or pilose, rarely glabrescent. |
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(34) |
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34 (33) |
Plants gray tomentose; terminal leaflet of radical leaves shortly petiolulate. |
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54 P. nervosa |
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Plants white tomentose; leaflets of radical leaves sessile. |
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(35) |
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35 (34) |
Leaflets abaxially densely silvery tomentose and appressed silvery sericeous, adaxially appressed silvery sericeous; flowers ca. 2 cm in diam.; petals purplish at base; stamens and styles purple. |
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56 P. argyrophylla |
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Leaflets abaxially densely gray or white tomentose, pilose on veins, adaxially green or gray-green, appressed sericeous or pilose; petals yellow; stamens and styles yellow-white. |
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(36) |
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36 (35) |
Leaflets adaxially appressed pilose, margin crenate; pedicels erect; flowers 1–1.4 cm in diam. |
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51 P. saundersiana |
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Leaflets adaxially appressed sericeous, margin lobed, lobes fasciated; pedicels nodding; flowers 1.5–2 cm in diam. |
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52 P. pendula |
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37 (24) |
Radical leaves pinnate. |
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69 P. supina |
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Radical leaves 3-foliolate, pedate, or palmate, rarely some leaves pinnate. |
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(38) |
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38 (37) |
Radical leaves palmately or pedately 5-foliolate, rarely some leaves 3-foliolate or pinnate. |
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(39) |
+ |
Radical leaves 3-foliolate. |
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(43) |
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39 (38) |
Stems prostrate, stoloniferous; roots slender; flowers 0.5–1 cm in diam. |
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68 P. kleiniana |
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Stems erect or decumbent; roots robust, usually woody; flowers 1.5–2.5 cm in diam. |
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(40) |
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40 (39) |
Plants glandular hairy. |
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(41) |
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Plants not glandular hairy. |
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(42) |
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41 (40) |
Plants red stipitate glandular hairy; radical leaves palmately or pedately 5-foliolate. |
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77 P. desertorum |
+ |
Plants sessile glandular hairy; radical leaves palmately 5-foliolate, some leaves 3-foliolate, rarely pinnate. |
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78 P. subdigitata |
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42 (40) |
Flowering stems and petioles pubescent, glabrescent; plants with leaf rosettes at anthesis. |
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75 P. chrysantha |
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Flowering stems and petioles white villous, hairs basally enlarged; plants without leaf rosettes at anthesis. |
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76 P. recta |
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43 (38) |
Flowering stems prostrate, elongate. |
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(44) |
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Flowering stems erect or ascending. |
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(46) |
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44 (43) |
Flowers many, in terminal cymes. |
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71 P. cryptotaeniae |
+ |
Flower solitary, lateral or terminal. |
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(45) |
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45 (44) |
Leaflets sessile, margin deeply serrate, stipules leathery, margin entire. |
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69 P. supina |
+ |
Leaflets markedly petiolulate, margin narrowly serrate, stipules membranous, margin entire or serrate. |
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70 P. centigrana |
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46 (43) |
Flowers 2.5–4 cm in diam. |
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73 P. macrosepala |
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Flowers not more than 2 cm in diam. |
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(47) |
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47 (46) |
Leaflets abaxially and sepals abaxially densely sessile glandular. |
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78 P. subdigitata |
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Leaflets abaxially and sepals abaxially not or only inconspicuously glandular. |
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(48) |
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48 (47) |
Plants spiny-setose, 10–20 cm tall. |
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72 P. asperrima |
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Plants not spiny-setose, more than 20 cm tall. |
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(49) |
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49 (48) |
Plants 50–100 cm tall; leaflets usually oblong or oblong-lanceolate, apex prolonged acuminate; stipules of cauline leaves lanceolate or oblong-lanceolate, mostly adnate to petiole. |
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71 P. cryptotaeniae |
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Plants 20–50 cm tall; leaflets usually elliptic, obovate, rhombic, or flabellate, apex obtuse or acute; stipules of cauline leaves broadly ovate or ovate-elliptic, free part longer than adnate part. |
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(50) |
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50 (49) |
Stems spreading; stipules of cauline leaves submembranous, margin entire or serrate; peduncle leafy; flowers 0.6–0.8 cm in diam. |
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70 P. centigrana |
+ |
Stems erect or ascending; stipules of cauline leaves herbaceous, margin entire; peduncle with minute, leaflike bracts; flowers 1–1.2 cm in diam. |
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74 P. xizangensis |