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FOC | Family List | FOC Vol. 5 | Urticaceae | Pilea

3. Pilea sect. Tetrameris C. J. Chen, Bull. Bot. Res., Harbin. 2(3): 44. 1982.

四萼组 si e zu

Subshrubs or perennial herbs. Leaves 3-veined. Male perianth lobes 4(or 5), valvate; female perianth lobes 4, subequal, shorter than achene.

About 13 species: SE Asia; 11 species (four endemic) in China.


1 Small subshrubs, less than 25 cm tall; stipules often over 20 mm, longitudinally 8-10-ribbed; inflorescences solitary, over 5 cm.   (2)
+ Herbs or subshrubs, often more than 30 cm tall; stipules less than 13 mm, longitudinally 2-ribbed; inflorescences in pairs, less than 4 cm.   (4)
       
2 (1) Leaf blade oblong-ovate, apex acuminate or shortly caudate; achene oblong-ovoid, spinulose-verrucose.   5 P. basicordata
+ Leaf blade ovate to suborbicular, apex obtuse, shortly acuminate, or cuspidate; achene broadly ovoid or subglobose, purplish spots or verrucose.   (3)
       
3 (2) Stems terete; leaf blade suborbicular; achene ca. 1.2 mm.   6 P. amplistipulata
+ Stems roundly 6-angled; leaf blade oblong or broadly ovate; achene ca. 2.8 mm.   7 P. hexagona
       
4 (1) Male flowers in clusters or forming pedunculate capitula.   (5)
+ Male flowers in dichotomous cymes, racemes, or panicles.   (7)
       
5 (4) Plants erect, less than 40 cm tall; leaf blade obovate, adaxial surface with 2, white grooves; male flowers forming pedunculate capitula, peduncle 1.5-4 cm.   15 P. cadierei
+ Plants erect, ascending, or climbing, usually over 50 cm tall; leaf blade lanceolate, uniformly colored, without adaxial white grooves; male flowers in axillary clusters.   (6)
       
6 (5) Leaf blade oblong- or linear-lanceolate, 6-10 × 1.5-2.8 cm; petiole 0.5-1.5 cm.   13 P. rubriflora
+ Leaf blade elliptic- or ovate-lanceolate, (3-)10-22 × (1.3-)4-6.5 cm; petiole (0.5-)2-7 cm.   14 P. funkikensis
       
7 (4) Leaf blade margin entire or erose, rarely inconspicuously serrate distally; achene with a purplish ring on both surfaces.   12 P. longicaulis
+ Leaf blade margin serrate or crenate-serrate; achene without colored ring on both surfaces.   (8)
       
8 (7) Male flowers in dichotomously branched cymes; leaf blade often lanceolate.   (9)
+ Male flowers in racemose cymes, few branched; leaf blade ovate or oblong.   (10)
       
9 (8) Leaf blade narrowly lanceolate, 7-15 × 2-4.5 cm; achene ovoid, spinulose-verrucose.   8 P. spinulosa
+ Leaf blade ovate to lanceolate, 2.5-10 × 1.2-3.5 cm; achene oblong-ovoid, tuberculate.   9 P. dolichocarpa
       
10 (8) Leaf blade oblong, apex acute; male glomerules scattered; achene ca. 2 mm, smooth, densely purplish spotted.   10 P. tsiangiana
+ Leaf blade ovate or ovate-lanceolate; male glomerules compact; achene ca. 1.2 mm, spinulose-verrucose.   11 P. pseudonotata

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