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FOC | Family List | FOC Vol. 8 | Crassulaceae | Sedum

2. Sedum sect. Oreades (Fröderström) K. T. Fu, Acta Phytotax. Sin. 12: 52. 1974.

山景天组 shan jing tian zu

Sedum "group" Oreades Fröderström, Acta Horti Gothob. 6(App.): 41. 1931.

Leaf base sessile, rarely subsessile, spurred. Inflorescence usually corymbiform, 1- to many flowered. Flowers pedicellate, very rarely subsessile. Sepals spurred or spurless. Petals yellow, rarely red or purple-red. Carpels erect to suberect, rarely divergent, not or only very slightly adaxially gibbous, base mostly broad and connate, less often free. Follicles erect or apically slightly divergent, not or only very slightly adaxially gibbous. Seeds mostly papillate.

About 67 species: Bhutan, China, India, Myanmar, Nepal, Pakistan, Sikkim; 64 species (54 endemic) in China.


1 Leaves verticillate or both opposite and alternate (all leaves alternate in S. chauveaudii var. margaritae).   (2)
+ Leaves alternate (sometimes opposite in S. blepharophyllum).   (8)
       
2 (1) Inflorescences cymose-scorpioid; flowers sessile or subsessile; sepals broadly ovate to suboblong, equal, ca. 1.5 mm   72 S. chuhsingense
+ Inflorescences corymbiform; flowers shortly pedicellate; sepals sublinear, unequal, 3-6.5 mm.   (3)
       
3 (2) Plants annual or biennial.   (4)
+ Plants perennial.   (5)
       
4 (3) Styles ca. 1.8 mm; nectar scales broadly linear-spatulate   70 S. leblancae
+ Styles ca. 0.5 mm; nectar scales quadrangular   71 S. tsiangii
       
5 (3) Leaves both verticillate and alternate; placenta subfalcate   66 S. bergeri
+ Leaves all verticillate (alternate in S. chauveaudii var. margaritae); placenta not subfalcate.   (6)
       
6 (5) Petals ± free, apex papillate   69 S. dielsii
+ Petals basally connate for more than 1.5 mm, apex not papillate.   (7)
       
7 (6) Sepals linear-spatulate; petals basally connate for ca. 1.5 mm   67 S. chauveaudii
+ Sepals linear-lanceolate; petals basally connate for 2-3 mm   68 S. tsonanum
       
8 (1) Seed funicle inflated at base, appendage stomach-shaped.   (9)
+ Seed funicle not inflated at base.   (16)
       
9 (8) Sepals spurless.   (10)
+ Sepals spurred.   (12)
       
10 (9) Seeds covered with thick, chaffy scales   58 S. ramentaceum
+ Seeds minutely papillate or hairy.   (11)
       
11 (10) Follicles 2- or 3-seeded; seeds minutely hairy   59 S. trichospermum
+ Follicles 12-14-seeded; seeds minutely papillate   60 S. prasinopetalum
       
12 (9) Leaves and sepals acuminate to cuspidate apically; petals 2-2.3 mm   65 S. erici-magnusii
+ Leaves and sepals obtuse to sometimes subacute apically; petals 4-7.2 mm.   (13)
       
13 (12) Sepals of terminal flower in 2 series   64 S. didymocalyx
+ Sepals in 1 series.   (14)
       
14 (13) Leaves oblong to subovate; nectar scales 1.6-1.8 mm   61 S. forrestii
+ Leaves linear-oblong, linear-lanceolate, or linear-obovate; nectar scales 0.4-0.7 mm.   (15)
       
15 (14) Leaves linear-lanceolate to linear-obovate; petals 4-6 mm; follicles 4- or 5-seeded   62 S. perrotii
+ Leaves linear-oblong; petals 7-7.2 mm; follicles usually ca. 7-seeded   63 S. longifuniculatum
       
16 (8) Placenta inflated, falcate to lunate.   (17)
+ Placenta not inflated, rarely falcate to lunate.   (31)
       
17 (16) Plants perennial; sterile stems present.   (18)
+ Plants annual or biennial; sterile stems absent.   (21)
       
18 (17) Flowers 5-merous; petals ca. 3.5 mm, apex obtuse   43 S. muyaicum
+ Flowers unequally 5-merous; petals 5.8-9 mm, apex acuminate.   (19)
       
19 (18) Underground stems absent; basal spur of leaves truncate-obtuse   46 S. longyanense
+ Underground stems present; basal spur of leaves obtuse or 3-lobed.   (20)
       
20 (19) Sepals 6.5-7.2 mm, spurless   44 S. celiae
+ Sepals 3.5-5 mm, spurred   45 S. dugueyi
       
21 (17) Sepals spurless at base.   (22)
+ Sepals spurred at base.   (25)
       
22 (21) Nectar scales broadly spatulate, ca. 0.4 mm, apically appendaged; leaf apex acuminate   47 S. celatum
+ Nectar scales narrowly linear to linear-spatulate, 0.7-1.3 mm, apically not appendaged; leaf apex obtuse.   (23)
       
23 (22) Nectar scales 0.8-1.3 mm; follicles 3-6-seeded; stamens 5   48 S. przewalskii
+ Nectar scales ca. 0.7 mm; follicles 1-3-seeded; stamens 10.   (24)
       
24 (23) Petals narrowed toward base, apex obscurely mucronate; placenta lunulate   49 S. pagetodes
+ Petals broadened toward base, apex not mucronate; placenta basal   50 S. pratoalpinum
       
25 (21) Leaves broadened in distal 1/2 (sometimes not so in S. sinoglaciale); nectar scales 1-1.5 mm.   (26)
+ Leaves broadened in proximal 1/2; nectar scales 0.5-0.8 mm.   (28)
       
26 (25) Follicles 2-seeded; antepetalous stamens inserted 1-1.2 mm from petal base; seeds smooth   57 S. semilunatum
+ Follicles 5- to many seeded; antepetalous stamens inserted ca. 2 mm from petal base; seeds papillate.   (27)
       
27 (26) Carpels basally connate for 1.2-1.8 mm; basal leaves spatulate, others subpanduriform; sepals obtuse at apex   55 S. chingtungense
+ Carpels ± free; leaves linear to linear-oblong; sepals acute at apex   56 S. sinoglaciale
       
28 (25) Nectar scales apically parted to lobate; leaf apex obtuse; follicles 2-4-seeded   54 S. purdomii
+ Nectar scales apically obtuse or retuse; leaf apex acute to acuminate; follicles (3-)7-12-seeded.   (29)
       
29 (28) Petals ca. 4 mm, ± free   53 S. fedtschenkoi
+ Petals 6-10.5 mm, basally subconnate.   (30)
       
30 (29) Petals yellow; placenta falcate; follicles 10-12-seeded   51 S. paracelatum
+ Petals purplish red; placenta sublunulate; follicles 7-9-seeded   52 S. luchuanicum
       
31 (16) Petals abruptly narrowed basally or attenuate into a claw.   (32)
+ Petals neither narrowed basally nor attenuate into claw.   (36)
       
32 (31) Plants annual or biennial; sterile stems absent.   (33)
+ Plants perennial; sterile stems present.   (34)
       
33 (32) Sepals lanceolate to lanceolate-oblong; petals red, obtrullate; stamens often 5   41 S. obtrullatum
+ Sepals ovate to ovate-oblong; petals yellow, narrowly oblong; stamens 10   42 S. raymondii
       
34 (32) Petal claw broad, less than 3 mm; sepals spurred, margin papillate   38 S. gagei
+ Petal claw narrow, to 4 mm; sepals spurless, margin laevigate.   (35)
       
35 (34) Petals sagittate, limb subovate, attenuate in distal 1/2, apex narrowly apiculate   39 S. sagittipetalum
+ Petals trullate, limb ovate-lanceolate to broadly ovate, basally narrowed, apex mucronate   40 S. trullipetalum
       
36 (31) Petals basally connate for more than 1 mm (for ca. 0.8 mm in S. costantinii); stamens in 2 series.   (37)
+ Petals free or basally subconnate usually for less than 1 mm; stamens in 1 or 2 series.   (42)
       
37 (36) Petal margin erose or suberose.   (38)
+ Petal margin entire.   (39)
       
38 (37) Petals lanceolate to oblong, margin conspicuously erose; leaves linear, apex spinose-apiculate; sepals spurless, apex spinose-apiculate   9 S. daigremontianum
+ Petals obovate-lanceolate to obovate, margin ± erose; leaves lanceolate to broadly oblong, apex acute; sepals spurred, apex acute   10 S. oreades
       
39 (37) Petals ca. 11 mm   12 S. feddei
+ Petals 5-6.2 mm   (40)
       
40 (39) Sepals yellowish green; leaf base spur 3-lobed   11 S. costantinii
+ Sepals green; leaf base spur entire.   (41)
       
41 (40) Petal base clawless, apex long mucronate; sepals linear or subspatulate   13 S. fui
+ Petal base ± clawed, apex subacuminate; sepals suboblong   14 S. wangii
       
42 (36) Leaf apex obtuse; stamens 5-10(-12).   (43)
+ Leaf apex acuminate to acute; stamens 10 (sometimes 5 in S. ulricae).   (52)
       
43 (42) Sepals spurless.   (44)
+ Sepals spurred.   (47)
       
44 (43) Flowers 5-merous; plants less than 2.5 cm tall.   (45)
+ Flowers unequally 5-merous; plants 3.5-6 cm tall.   (46)
       
45 (44) Plants 1-2.5 cm tall; leaves narrowly oblong; follicles 6-8(-14)-seeded; seeds minutely papillate   28 S. fischeri
+ Plants ca. 10 mm tall; leaves ovate to suborbicular; follicles 1- or 2-seeded; seeds winged at both ends   29 S. oligocarpum
       
46 (44) Leaves broadly linear to oblong; follicles few seeded   30 S. lutzii
+ Leaves ovate; follicles 6- or 7-seeded   31 S. tsinghaicum
       
47 (43) Seeds smooth, winged; leaf and sepal margin smooth; follicles 10-12-seeded   37 S. magniflorum
+ Seeds papillate, wingless; leaf and sepal margin papillate; follicles more than 12-seeded.   (48)
       
48 (47) Petals basally free or almost so.   (49)
+ Petal basally subconnate.   (50)
       
49 (48) Petal apex smooth; nectar scales apically emarginate; follicles 12-15-seeded   32 S. roborowskii
+ Petal apex papillate; nectar scales apically rounded; follicles 20-22-seeded   33 S. blepharophyllum
       
50 (48) Leaves broadly lanceolate to lanceolate-oblong; petal apex papillate   34 S. obtusipetalum
+ Leaves obovate, obovate-oblong, or suboblong; petal apex smooth.   (51)
       
51 (50) Petals broadly triangular; sepal apex rounded; stamens 5 or 10   35 S. somenii
+ Petals narrowly oblong; sepal apex obtuse; stamens 10   36 S. morotii
       
52 (42) Plants perennial; sterile stems numerous.   (53)
+ Plants annual or biennial; sterile stems absent (present in S. susanneae).   (58)
       
53 (52) Leaf base spur obtuse or truncate.   (54)
+ Leaf base spur lobed to sublobed.   (55)
       
54 (53) Sepals spurless; petals basally almost free; basal leaves of flowering stems ovate-lanceolate, persistent; seeds winged   19 S. barbeyi
+ Sepals spurred; petals basally connate for ca. 1.2 mm; basal leaves of flowering stems linear, soon withered; seeds wingless   20 S. pampaninii
       
55 (53) Spur 2-lobed, occasionally ± 3-lobed; leaf and sepal margin glandular ciliate   18 S. glaebosum
+ Spur 3-lobed or ± so (sometimes truncate in S. heckelii); leaf and sepal margin glabrous.   (56)
       
56 (55) Sepals spurless, adaxially papillate   15 S. beauverdii
+ Sepals spurred, adaxially smooth.   (57)
       
57 (56) Inflorescences dense; sepals narrowly oblong to ovate, apex shortly acuminate   16 S. heckelii
+ Inflorescences lax; sepals lanceolate to narrowly triangular, apex acuminate   17 S. nothodugueyi
       
58 (52) Sepals spurless.   (59)
+ Sepals spurred.   (60)
       
59 (58) Leaf and sepal margin glabrous; carpels glabrous   21 S. susanneae
+ Leaf and sepal margin spinescent ciliate; carpels adaxially spinescent hairy   22 S. stimulosum
       
60 (58) Petals 6.5-8.9 mm.   (61)
+ Petals less than 4.5 mm.   (62)
       
61 (60) Pedicel 1-2 mm; sepals linear-lanceolate or subovate; petals apex long mucronate   23 S. rosei
+ Pedicel 6-10 mm; sepals oblong; petal apex acuminate   24 S. platysepalum
       
62 (60) Petals ca. 3 mm, basally connate, apex mucronulate   27 S. henrici-robertii
+ Petals to 4.5 mm, free, apex not mucronulate.   (63)
       
63 (62) Plants 10-13 cm tall; carpels suberect   25 S. franchetii
+ Plants 3-6 cm tall; carpels divergent   26 S. ulricae

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