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FOC | Family List | FOC Vol. 8 | Saxifragaceae | Saxifraga

4. Saxifraga sect. Ciliatae Haworth, Misc. Nat. 160. 1803.

山羊臭组 shan yang xiu zu

Hirculus Haworth.

Herbs mostly perennial, some possibly annual, biennial, or monocarpic perennial (suspected rather than demonstrated), forming erect individuals, cespitose clumps, cushions, or sprawling mats, lacking basal bulbils. Leaves alternate, very rarely opposite, petiolate or not; leaf blade herbaceous or carnose, lacking chalk glands and crystals, margin usually entire, occasionally toothed or lobed. Flowering stem usually leafy; inflorescence cymose or flower solitary. Flowers sometimes unisexual, actinomorphic, 5-merous. Petals usually yellow or orange, occasionally white, pink, or red, sometimes spotted orange, red, or purple, very often with callosities. Filaments subulate to linear; pollen 3-nucleate, exine tectum finely striate. Carpels connate for most of placental region, dehiscing distally in fruit; ovary subsuperior to subinferior. Integuments 2.

About 175 species: Asia, Europe, North America, mainly in arctic and montane regions; 166 species (112 endemic) in China.


Key 4

1 Basal leaves pubescent adaxially.   (2)
+ Basal leaves glabrous adaxially.   (7)
       
2 (1) Hairs on adaxial surface of basal leaves setose or eglandular   158 S. gyalana
+ Hairs on adaxial surface of basal leaves glandular.   (3)
       
3 (2) Basal or proximal cauline leaves with at least 3 forward-pointing, triangular lobes.   (4)
+ Basal and cauline leaves entire.   (5)
       
4 (3) Basal leaves with at least 3 forward-pointing, triangular lobes   173 S. candelabrum
+ Basal leaves entire   172 S. dielsiana
       
5 (3) Flowering stem leafless, bearing bracts only   169 S. lixianensis
+ Flowering stem leafy.   (6)
       
6 (5) Petals yellow; plants often forming a mat of basal leaf rosettes   170 S. sediformis
+ Petals abaxially purple, adaxially lilac near base and yellow near apex; plants usually with a single basal leaf rosette   171 S. daochengensis
       
7 (1) Basal leaves adaxially pustulate distally.   (8)
+ Basal leaves adaxially smooth (or wrinkled on drying).   (11)
       
8 (7) Inflorescence 2- or 3-flowered or flower solitary; petals pale yellow to yellow or ivory   163 S. punctulata
+ Inflorescence 2-10-flowered; petals yellow or white.   (9)
       
9 (8) Margin of basal leaves entire   166 S. umbellulata
+ Margin of basal leaves setose-ciliate.   (10)
       
10 (9) Petals white, purple spotted, to 7 mm   164 S. punctulatoides
+ Petals yellow, purple spotted, more than 7 mm   165 S. yushuensis
       
11 (7) Margin of basal leaves entire.   (12)
+ Margin of basal leaves setose- or denticulate-ciliate.   (13)
       
12 (11) Petals yellow, orange spotted; inflorescence cymose   155 S. unguiculata
+ Petals yellow, unspotted; inflorescence subumbellate   166 S. umbellulata
       
13 (11) Petals red, pink, or white on both surfaces or only abaxially.   (14)
+ Petals yellow on both surfaces.   (16)
       
14 (13) Petals more than 7 × 2.5 mm   166 S. umbellulata
+ Petals less than 7 × 2.5 mm.   (15)
       
15 (14) Petals deep red abaxially or on both surfaces   167 S. sanguinea
+ Petals white or pinkish on both surfaces   161 S. signatella
       
16 (13) Basal leaf rosettes usually at least 2 cm in diam.   (17)
+ Basal leaf rosettes rarely to 2 cm in diam.   (18)
       
17 (16) Petals unspotted   166 S. umbellulata
+ Petals adaxially purple spotted proximally   168 S. signata
       
18 (16) Longest cauline leaves at least 7 mm.   (19)
+ Longest cauline leaves less than 7 mm.   (21)
       
19 (18) Sepals glabrous   157 S. taraktophylla
+ Sepals glandular pilose abaxially and marginally.   (20)
       
20 (19) Proximal internodes ca. 2.5 mm   155 S. unguiculata
+ Proximal internodes 3.5-4 mm   156 S. pellucida
       
21 (18) Pedicels glabrous; petals less than 3.5 mm   160 S. elatinoides
+ Pedicels (e)glandular pilose, at least sparsely so; petals more than 4 mm.   (22)
       
22 (21) Sepals spreading at anthesis   162 S. brunneopunctata
+ Sepals reflexed at anthesis.   (23)
       
23 (22) Cauline leaves markedly overlapping   155 S. unguiculata
+ Cauline leaves remote, scarcely, if at all, overlapping   159 S. heterotricha

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