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FOC | Family List | FOC Vol. 23 | Cyperaceae

37. Carex sect. Physocarpae Drejer ex L. H. Bailey in J. M. Coulter, Man. Bot. Rocky Mt. 382. 1885.

胀囊薹草组 zhang nang tai cao zu

Authors: Lun-Kai Dai, Tetsuo Koyama & Gordon C. Tucker

Culms usually rather tall and slightly stout. Leaf blades with transverse septate nodes. Involucral bracts leaflike, longer than inflorescence, usually not sheathed, rarely shortly sheathed. Spikes unisexual, several, few species with male flowers at top of female spike or with female flowers at top of male spike; 1-4 spikes at top of culm male, narrowly cylindric; remaining spikes female, single in involucral bract axil, oblong or cylindric, densely many flowered, usually shortly pedunculate. Female glumes oblong-lanceolate or narrowly ovate, muticous. Utricles obliquely patent, later rarely horizontally divaricate or deflexed, broadly ovate or ovate, inflated trigonous, membranous or subleathery, glabrous, distinctly veined, apex abruptly contracted into a short to long beak, orifice shortly 2-toothed or indistinctly toothed. Nutlets loosely enveloped in utricle, broadly obovate, trigonous; style slender, usually tortuous; stigmas 3.

About 20 species: E and SE Asia, Europe, North America; ten species in China.

1 Utricles ovate or broadly so, 6-12 mm   (2)
+ Utricles oblong-ovate or ovate, 4-5.5 mm   (5)
2 (1) Female spikes cylindric, 3-7.5 cm; utricles 6-8 mm   (3)
+ Female spikes oblong or ovoid, 1.5-3 cm; utricles 9-12 mm   (4)
3 (2) Leaf blades 8-15 mm wide; male spikes usually 3-7, sometimes with few female flowers at base, with few male flowers at top of female spike; female glumes oblong-lanceolate; utricles horizontally divaricate in maturity, ovoid, apex abruptly contracted into a slightly long beak.   346 C. rhynchophysa
+ Leaf blades 2-5 mm wide; male spikes 2 or 3, spikes unisexual; female glumes narrowly lanceolate; utricles obliquely patent in maturity, oblong-ovate, apex attenuate into a rather short beak.   347 C. vesicaria
4 (2) Male spike 1; female spikes ovoid, 1.5-2 × ca. 1.5 cm, distance approximate, usually congregated at top of culm; utricles broadly ovate; nutlets broadly rhomboid, style base slightly thickened.   348 C. dickinsii
+ Male spikes 1 or 2; female spikes oblong, 2-3 × 1-1.2 cm, distance remote; utricles oblong-ovate; nutlets obovate, style thickened at lower half.   349 C. idzuroei
5 (1) Male spike 1, terminal; utricles with recurved and slightly long stipe.   350 C. mollissima
+ Male spikes 2 to several, inserted at top of culm; utricles estipitate   (6)
6 (5) Utricles oblong-ovate, apex attenuate into a beak, emarginate at orifice, not 2-toothed   (7)
+ Utricles ovate or broadly ovate, apex abruptly narrowed into a beak, slightly long or rather shortly 2-toothed at orifice   (8)
7 (6) Culms 60-90 cm tall; female glumes oblong-lanceolate, slightly shorter than or nearly equaling utricle; utricles 5-5.5 mm.   351 C. pamirensis
+ Culms 15-50 cm tall; female glumes ovate, ca. 1/2 length of utricle or slightly longer than utricle; utricles 4-4.5 mm.   352 C. dichroa
8 (6) Female glumes oblong-lanceolate, slightly shorter than or nearly equaling utricle; utricles ca. 4 mm.   353 C. rostrata
+ Female glumes ovate or narrowly lanceolate, shorter than utricle; utricles ca. 5 mm   (9)
9 (8) Female glumes narrowly lanceolate, brown-purple.   354 C. obscuriceps
+ Female glumes ovate, brown-yellow.   355 C. vesicata

Lower Taxa


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