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FOC | Family List | FOC Vol. 23 | Cyperaceae

34. Carex sect. Rhomboidales Kükenthal in Engler, Pflanzenr. 38(IV. 20): 622. 1909.

菱形果薹草组 ling xing guo tai cao zu

Authors: Prof. Song-Yun Liang, Tetsuo Koyama & Gordon C. Tucker

Culms central or lateral, rarely very short. Involucral bracts shortly bladed and sheathing or leaflike and sheathless. Terminal spike male, cylindric, clavate, or linear; lateral spikes androgynous, usually with a long or short male part at apex, rarely all female. Female glumes ovate, oblong, or ovate-lanceolate, rarely linear-lanceolate, apex mucronate, rounded or acuminate, rarely emarginate. Utricles rhomboid, elliptic, or ovate, subleathery or membranous, glabrous, rarely hirsute, many veined, base gradually attenuate, apex abruptly contracted into a long beak, orifice 2-toothed. Nutlets obovate or ovate, trigonous, with angles constricted at middle, beak erect or curved, annulate or not at orifice; style base thickened or not thickened; stigmas 3.

About 50 species: E and SE Asia; 43 species (34 endemic) in China.


1 Culms very short   (2)
+ Culms long   (3)
       
2 (1) Female glumes ovate, rounded at apex.   331 C. radiciflora
+ Female glumes linear-lanceolate, acuminate at apex, scabrous on margins.   332 C. zunyiensis
       
3 (1) Nutlets trigonous with angles not constricted at middle   (4)
+ Nutlets trigonous with angles constricted at middle   (18)
       
4 (3) Utricles hirsute   (5)
+ Utricles glabrous   (7)
       
5 (4) Female spikes usually bearing male flowers at base; utricles rhomboid or obovate.   290 C. chinensis
+ Female spikes without male flowers at base; utricles ovate or lanceolate-ovate   (6)
       
6 (5) Utricles lanceolate-ovate, 4.5-5 mm.   291 C. diplodon
+ Utricles ovate, 3-4 mm.   292 C. blinii
       
7 (4) Nutlets with transparent grains on surfaces.   304 C. oxyphylla
+ Nutlets without transparent grains on surfaces   (8)
       
8 (7) Style slender, base curved.   293 C. qingdaoensis
+ Style base slightly thickened, not curved   (9)
       
9 (8) Involucral bracts leaflike, all involucral bracts or lowest one longer than inflorescence   (10)
+ Involucral bracts shortly bladed, shorter than inflorescence   (12)
       
10 (9) Spikes 4 or 5, terminal spike linear, sessile.   294 C. manciformis
+ Spikes 2-4, terminal spike linear-cylindric, linear-oblong, or linear, pedunculate   (11)
       
11 (10) Spikes 3 or 4, terminal spike male, linear-cylindric, lateral spikes androgynous, male part 1/3-1/2 length of female part.   297 C. austrosinensis
+ Spikes 2-4, terminal spike male, linear or linear-oblong, lateral spikes female, oblong.   298 C. macrandrolepis
       
12 (9) Utricles hirsute   (13)
+ Utricles glabrous   (14)
       
13 (12) Utricles narrowly rhomboid, 5-6 mm.   299 C. tatsutakensis
+ Utricles oblong, 7-8 mm.   300 C. longerostrata
       
14 (12) Leaves 6-10 mm wide   (15)
+ Leaves 1.5-3.5 mm wide   (16)
       
15 (14) Leaves subequaling culm, 6-8 mm wide; nutlets with beak erect, base erect stipitate.   301 C. xiphium
+ Leaves surpassing culm, 7-10 mm wide; nutlets with beak curved, base curved stipitate.   303 C. yangshuoensis
       
16 (14) Spikes 2, terminal spike clavate, ca. 1 cm, lateral spikes female, ovate, ca. 1 cm, 4- or 5-flowered.   302 C. pseudolongerostrata
+ Spikes 3 or 4, terminal spike cylindric, 3-6 cm, lateral spikes mostly female, ± with male flowers at apex, cylindric, 1.5-4.5 cm, slightly densely flowered   (17)
       
17 (16) Leaves 3-5 mm wide; utricle beak 2-toothed; nutlets ovoid, ca. 5 mm.   295 C. simulans
+ Leaves 7-12 mm wide; utricle beak barely 2-toothed; nutlets broadly obovoid, 3.5-4.5 mm.   296 C. austrozhejiangensis
       
18 (3) Culms lateral   (19)
+ Culms central   (31)
       
19 (18) Nutlets with curved beak   (20)
+ Nutlets with erect beak   (25)
       
20 (19) Nutlets with longer beak, ca. 1 mm; female spikes oblong, 1.7-3 cm   (21)
+ Nutlets with shorter beak; female spikes cylindric, 3-7 cm   (22)
       
21 (20) Leaf blades, involucral bracts, and sheaths scabrous; involucral bracts leaflike, longer than spike; female glumes pale brown, green 3-veined scabrous costa excurrent into a long awn at obtuse apex.   305 C. shangchengensis
+ Leaf blades, involucral bracts, and sheaths glabrous; involucral bracts shortly bladed, shorter than spike; female glumes yellow-white, green 3-veined glabrous costa excurrent into a mucro at emarginate apex.   306 C. manca
       
22 (20) Female spikes rather thick, ca. 15 mm wide; utricles subglobose.   307 C. nanchuanensis
+ Female spikes rather slender, 7-11 mm wide; utricles ovate   (23)
       
23 (22) Lateral spikes androgynous, usually male part equaling female part, densely flowered.   310 C. thibetica
+ Lateral spikes mostly female with male flowers at apex, male part shorter than female part   (24)
       
24 (23) Female spikes sparsely flowered, horizontally patent at maturity.   308 C. heudesii
+ Female spikes densely flowered, obliquely patent at maturity.   309 C. taipaishanica
       
25 (19) Female spikes only 1; female glumes ovate, costa projecting beyond apex forming a long awn.   311 C. funingensis
+ Female spikes 3-5; female glumes ovate or ovate-oblong, obtuse or rounded at apex, green 3-veined costa projecting beyond apex forming a mucro or cusp but rarely an awn   (26)
       
26 (25) Female spikes cylindric; nutlets with cylindric beak annulate at orifice   (27)
+ Female spikes ovate or oblong; beak of nutlets not thickened or slightly thickened but not annulate at orifice   (29)
       
27 (26) Female spikes spaced, sparsely flowered; female glumes with a mucro at apex; utricles puberulent.   314 C. longqishanensis
+ Female spikes approximate, densely flowered; female glumes awned or cuspidate at apex; utricles glabrous   (28)
       
28 (27) Female glumes awned at apex; terminal spike male, linear-cylindric, 4.5-8 cm.   312 C. harlandii
+ Female glumes cuspidate at apex; terminal spike male, linear 1.5-3 cm.   313 C. hoozanensis
       
29 (26) Utricles puberulent.   306 C. manca
+ Utricles glabrous   (30)
       
30 (29) Culms 25-40 cm tall; leaf blades 4-6 mm wide.   315 C. hastata
+ Culms 7-13 cm tall; leaf blades 10-15 mm wide.   316 C. kaoi
       
31 (18) Culms, leaves, involucral bracts, female glumes, and utricles all puberulent   (32)
+ Culms, leaves, involucral bracts, and female glumes all glabrous   (33)
       
32 (31) Beak of nutlets erect; female spikes 3, bearing a short male part at apex; leaf blades 6-10 mm wide.   317 C. pseudolaticeps
+ Beak of nutlets curved; female spikes 1 or 2; leaf blades 3-5 mm wide.   318 C. laticeps
       
33 (31) Female glumes linear-lanceolate or ovate-lanceolate, usually acuminate at apex, rarely awned   (34)
+ Female glumes ovate, oblong-ovate, or oblong, obtuse, rounded, or truncate at apex, green 1- or 3-veined costa projecting beyond apex forming a mucro or long awn   (38)
       
34 (33) Female glumes brown   (35)
+ Female glumes pale green or pale yellow   (36)
       
35 (34) Female glumes linear-lanceolate; lateral spikes usually female, occasionally becoming androgynous, male part shorter than female part; style base not thickened.   329 C. brevicuspis
+ Female glumes ovate-lanceolate; lateral spikes usually female, occasionally becoming androgynous, male part longer than or equaling female part; style base thickened.   330 C. kuchunensis
       
36 (34) Beak of nutlets curved; female glumes slightly longer than or equaling utricle.   326 C. cheniana
+ Beak of nutlets erect; female glumes shorter than utricle   (37)
       
37 (36) Culms up to ca. 65 cm tall; leaf blades 8-10 mm wide; lateral spikes cylindric, ca. 6.5 × 1 cm.   327 C. saxicola
+ Culms 2-10 cm tall; leaf blades 1.5-2.5 mm wide; lateral spikes oblong-cylindric, ca. 1.2 × 0.6 cm.   328 C. lianchengensis
       
38 (33) Beak of nutlets short and curved, annulate at orifice, style base thickened; leaf blades abruptly contracted into a cusp at apex.   319 C. liqingii
+ Beak of nutlets long and erect, rarely short, annulate or not at orifice, style base hardly thickened; leaf blades not contracted into a cusp at apex   (39)
       
39 (38) Nutlets shallowly constricted on angles at middle, beak short; leaf blades papery.   320 C. rhynchophora
+ Nutlets deeply constricted on angles at middle, beak ca. 1 mm or more; leaf blades leathery   (40)
       
40 (39) Female glumes ovate, acute or mucronate at apex, red-brown; nutlets constricted on angles and concave surfaced.   321 C. calcicola
+ Female glumes oblong, obtuse or emarginate at apex, green 3-veined costa excurrent into an awn, brown; nutlets constricted only on angles at middle   (41)
       
41 (40) Spikes 2, lateral spike wholly female.   325 C. putuoensis
+ Spikes 3 or 4, lateral spikes female or androgynous with male part up to 1/2 length of spike   (42)
       
42 (41) Nutlets rhomboid.   322 C. tenebrosa
+ Nutlets obovoid or ovoid-ellipsoid   (43)
       
43 (42) Nutlets ovoid-ellipsoid; female glumes with awn ca. 3.5 mm.   323 C. chaofangii
+ Nutlets broadly obovoid; female glumes with awn ca. 5 mm.   324 C. wahuensis

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