All Floras      Advanced Search
FOC Vol. 23 Page 302, 378, 382 Login | eFloras Home | Help
FOC | Family List | FOC Vol. 23 | Cyperaceae

32. Carex sect. Molliculae Ohwi, Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 11: 450. 1936.

柔果薹草组 rou guo tai cao zu

Authors: Lun-Kai Dai, Tetsuo Koyama & Gordon C. Tucker

Rhizome stoloniferous. Involucral bracts not sheathed or shortly sheathed. Spikes usually unisexual, terminal spike male, lateral spikes female (Carex harrysmithii and C. agglomerata with unisexual spikes and bisexual spikes), cylindric, rarely oblong, densely many flowered. Female glumes ovate or lanceolate-oblong, apex acuminate, mucronate, aristate, or muticous. Utricles yellowish green or stramineous, erect or obliquely patent, ovate, elliptic, or oblong, slightly inflated or inflated trigonous, many veined, beak medium sized to long or rather long, orifice shortly 2-toothed, rarely obliquely truncate or emarginate. Nutlets rather loosely enveloped; stigmas 3.

About 20 species: E and SE Asia; 18 species (nine endemic) in China.

1 Spikes unisexual and bisexual, uppermost 1 or 2 spikes androgynous or gynaecandrous, remaining spikes female, or uppermost 1 or 2 spikes male and remaining female or bisexual   (2)
+ Spikes usually unisexual, uppermost 1 or 2 spikes male (C. planiculmis sometimes bearing male spikes with several female flowers at top), remaining spikes female, sometimes with few male flowers at top (e.g., C. doniana)   (6)
2 (1) Uppermost 1 or 2 spikes bisexual, androgynous or gynaecandrous, remaining spikes female, spikes oblong, not over 2 cm, approximate, usually congregate at top of culm; female glumes stramineous or yellowish green   (3)
+ Uppermost 1 or 2 spikes male, remaining spikes female or bisexual, bisexual spikes androgynous or gynaecandrous, spikes cylindric, 2.5-10 cm, remote, not congregate at top of culm; female glumes purple or brown-red   (4)
3 (2) Spikes 2, rarely 3, androgynous, with few male flowers at top of spike; female glumes lanceolate, ca. 2 mm; utricles ca. 3 mm; nutlets stipitate.   271 C. harrysmithii
+ Spikes 3 or 4, rarely 2, terminal spike gynaecandrous, lateral spikes female; female glumes ovate, ca. 3 mm; utricles 3.5-4 mm; nutlets sessile.   272 C. agglomerata
4 (2) Rhizome stoloniferous; leaves glabrous; female spikes densely many flowered, usually with few male flowers at base; female glumes muticous or mucronate; utricles with a short beak.   273 C. alliiformis
+ Rhizome without stolons; leaves pubescent abaxially; female spikes laxly flowered, usually with few male flowers at top; female glumes long aristate or awned; utricles with a long beak   (5)
5 (4) Leaves 7-12 mm wide; male spike 7-10 cm; utricles spreading.   274 C. purpureotincta
+ Leaves up to 17 mm wide; male spike 4-6 cm; utricles reflexed.   275 C. xiangxiensis
6 (1) Utricles wholly scabrid pubescent.   282 C. scabrisacca
+ Utricles glabrous, not scabrid   (7)
7 (6) Spikes approximate, usually congregate at top of culm, male spikes clavate, 1.5-2 cm, female spikes oblong, not over 2.5 cm (only C. huashanica with male spikes 2-5 cm, female spikes 3-6 cm), subsessile or shortly pedunculate   (8)
+ Spikes remote, not congregate at top of culm, male spike and female spikes cylindric (C. japonica with spikes oblong), 2.5-7 cm, lower spikes long pedunculate, upper spikes shortly pedunculate   (12)
8 (7) Female spikes cylindric, 3-6 cm; female glumes with long awn at apex, scabrous on margins at awn; utricles ca. 5 mm, obliquely truncate or emarginate at orifice.   276 C. huashanica
+ Female spikes oblong, not over 2.5 cm; female glumes muticous or mucronate; utricles 3-4 mm, 2-toothed at orifice (except in C. submollicula)   (9)
9 (8) Leaf blades 2-4 mm wide; utricles ca. 5 mm, distinctly veined.   280 C. submollicula
+ Leaf blades 4-10 mm wide; utricles 3-4 mm, obscurely veined   (10)
10 (9) Female spikes 1.5-2.5 cm; utricles oblong-ovate, slightly inflated trigonous, membranous, usually rugulose, apex abruptly contracted into a medium-sized to long beak.   279 C. mollicula
+ Female spikes 0.8-1.7 cm; utricles elliptic, inflated trigonous, thickly papery, smooth, apex abruptly contracted into a short beak   (11)
11 (10) Female glumes ovate or narrowly ovate; nutlets obovate.   277 C. aphanolepis
+ Female glumes oblong; nutlets subrhomboid.   278 C. radicina
12 (7) Leaf blades 5-12 mm wide; female spikes (1.5-)2-7 cm; utricles attenuate to a short beak at apex, nearly obliquely truncate or obscurely 2-toothed at orifice   (13)
+ Leaf blades 2-5 mm wide; female spikes 1-3(-4) cm; utricles attenuate to medium-sized or long beak at apex, distinctly 2-toothed at orifice   (15)
13 (12) Culms compressed triquetrous, one angle obscure; male spike not over 3.5 cm, with female flowers at top, female spikes 1.5-4.5 cm, without male flowers at top; female glumes ovate, muticous or sometimes mucronate at apex; utricles ovate, with a recurved beak, style deciduous, stigmas short.   288 C. planiculmis
+ Culms triquetrous; male spike 3-7.5 cm, female spikes 2-7 cm, sometimes with several male flowers; female glumes ovate-lanceolate, mucronate at apex; utricles oblong-ovate, with a straight beak; style usually persistent, stigmas rather long   (14)
14 (13) Spikes 2-6, male spike linear-cylindric, 3-7.5 cm, female spikes cylindric, 3-7 cm; female glumes pale yellow; utricles 3.5-4 mm, greenish yellow; basal leaf sheaths brownish.   286 C. doniana
+ Spikes 3 or 4, male spike narrowly cylindric, ca. 3 cm, female spikes cylindric, 2-3.5 cm; female glumes yellow-brown; utricles 4-4.5 mm, olivaceous; basal leaf sheaths purple-brown.   287 C. phaenocarpa
15 (12) Female spikes oblong or oblong-cylindric, 1-2.5 cm; utricles 4-5 mm, stramineous, slightly nitid, obscurely veined.   281 C. japonica
+ Female spikes cylindric, to 3 cm or more; utricles ca. 3.5 mm, yellowish green, not nitid, distinctly veined   (16)
16 (15) Culms 30-60 cm tall; male spike clavate, 2-3 cm; female glumes narrowly ovate, muticous at apex; utricles ovate, slightly inflated.   285 C. alopecuroides
+ Culms not over 30 cm tall; male spike linear, 1-5 cm; female glumes oblong-ovate, mucronate at apex; utricles elliptic or oblong, not inflated   (17)
17 (16) Utricles elliptic, 3-3.5 mm, with a medium-sized to long beak, beak ca. 1/3 length of utricle.   283 C. subtransversa
+ Utricles oblong, 4-5 mm, with a long beak, beak ca. 1/2 length of utricle.   284 C. liui

  • List of lower taxa


  • Flora of China @
    Browse by
    Advanced Search

    Flora of China Home




     |  eFlora Home |  People Search  |  Help  |  ActKey  |  Hu Cards  |  Glossary  |