Plants terrestrial, medium-sized. Rhizome creeping, forked, sparsely scaly; scales shiny, lanceolate or ovate-lanceolate, margin with sparse glandular-hairlike teeth, apex long acuminate. Fronds approximate; stipe chestnut-brown or stramineous, nearly as long as lamina, with scales, hyaline multicellular hairs, and scalelike hairs, grooved adaxially; lamina 2-pinnate to 3-pinnate-pinnatifid, broadly ovate to ovate-lanceolate, thinly herbaceous, not narrowed to base, apex subacute or acuminate; pinnae mostly subopposite, spreading or ascending, pinnatisect to 2-pinnate, broadly lanceolate to lanceolate, slightly falcate at apex, equilateral, truncate or rounded-cuneate at base, subsessile or very shortly stalked, apex acuminate; basal pair of pinnae not shortened; pinnules sessile, alternate, only basal pair of pinnules subopposite, slightly shorter than upper pinnules, ovate-lanceolate or lanceolate, equilateral, base subtruncate or rounded-cuneate, basiscopic base slightly wider than acroscopic base, apex obtuse or acuminate; pinnule lobes oblong, sessile or adnate to costae or costules, lobed or parted, obtuse. Veins visible but not clearly so, free, pinnate, simple or forked, reaching marginal teeth of lobes. Lamina surface, all costae, and veins with hyaline long multicellular hairs and narrow scalelike hairs. Sori small, orbicular, abaxial on veins, sorophore protuberant; sporangia with slender stalks, annulus cells 12-14; indusia small, laterally equilateral, pale green, broadly ovate, membranous, glandular hairy, sparsely ciliate at margin, attached proximally to sorophore, hidden by sporangia at maturity; spores yellow, bean-shaped or narrowly spheroidal, perispore with dense inflated, bulbous processes. x = 42.

Three species: tropical to temperate areas of SE and E Asia and New Zealand; three species (one endemic) in China.

(Authors: Wang Zhongren (王中仁); Masahiro Kato)