Rhizome bearing scales and hairs or only scales; roots borne on all sides of rhizome. Scales glabrous or with multiseptate hairs, basifixed, base not cordate. Stipe articulate at base to phyllopodia, grooved or not, glabrous; lamina compound, tripinnate or quadripinnate (rarely in small plants bipinnate) toward base and in middle part, deltoid and broadest toward base, glabrous (sometimes minute hairs present), not or slightly dimorphic (often fertile fronds more strongly dissected); pinnae deltoid or narrowly triangular; pinnules of at least larger pinnae anadromous; pinnules or pinna lobes narrowly ovate. Leaf axes glabrous (sometimes a few minute hairs present). Veins in ultimate lobes simple, not reaching margin; false veins not present. Sori indusiate, frequently single on a segment, terminal on veins; indusium scalelike, either attached at narrow, cordate base only, or attached at base and only part of sides, very long stalked, 3-seriate; annulus longitudinal, consisting of ca. 16 thickened cells. Spores elliptic, monolete, closely tuberculate, translucent. x = 41.

Two species: S and SE Asia to Pacific islands; one species in China.

(Authors: Xing Fuwu (邢福武), Wang Faguo (王发国); Hans P. Nooteboom)