Crypsinus C. Presl; Phymatopsis J. Smith (1875), not Tulasne ex Treviranus (1857); Phymatopteris Pichi Sermolli; Pichisermollia Fraser-Jenkins (2008), not H. C. Monteiro (1980); Pichisermollodes Fraser-Jenkins & Challis.

Rhizome slender, woody, densely scaly; scales reddish brown, brown, dark brown, or nearly black, rarely whitish, opaque, ovate-lanceolate or lanceolate, margin entire, toothed, or ciliate, apex attenuate or setaceous. Fronds remote, articulate to rhizome, monomorphic or dimorphic with longer and narrower fertile fronds. Stipe usually glabrous, rarely pubescent, scaly at base. Lamina simple, hastately 3-lobed, palmately or pinnately divided, sometimes pinnatisect with adnate pinnae, herbaceous or leathery, glabrous or pubescent, rarely scaly on abaxial side. Lobes usually lanceolate, margin distinctly cartilaginous, entire, notched, or serrate, apex acuminate or obtuse. Lateral veins (in lamina or segments) distinct; veinlets anastomosing to form areoles with included free veinlets. Sori orbicular, in 1 row on either side of costa, less often linear or in many rows, usually superficial, rarely sunken on abaxial surface and raised on adaxial surface. Sporangia with long stalk, annulus with 14 hardened cells. Spores brown, ellipsoid, surface tuberculate or verrucate. x = 12, n = 36, 37.

Until relatively recently, most species of Selliguea were included within Phymatopteris, but the concept widely used for Phymatopteris clearly includes the type species of Crypsinus, C. pyrolifolius (Goldmann) Copeland, which is very similar to S. rhynchophylla in this account. Both Smith et al. (Taxon 55: 719. 2006) and Fraser-Jenkins (Taxon. Revis. Indian Subcontinental Pteridophytes, 44. 2008) do not recognize Phymatopteris and include Crypsinus within Selliguea. Christenhusz et al. (Phytotaxa 19: 33. 2011) maintain Phymatopteris but include Crypsinus within Selliguea, a position that does not seem tenable in the light of the above observation.

Fraser-Jenkins erected the genus Pichisermollodes for those species with Polypodium-like pinnatifid to pinnate fronds with notched and toothed margins, but this genus was not accepted by Christenhusz et al.

About 75 species: tropical and subtropical Asia, Australia, Pacific islands, South Africa, and Madagascar; 48 species (29 endemic) in China.

(Authors: Lu Shugang (陆树刚); Peter H. Hovenkamp, Michael G. Gilbert)