aerophores beneath pinna bases swollen; segments 25-40 pairs, rectangular, 3-6 × 2-4 mm, margins usually dentate, apices truncate or rounded-truncate; veinlets 4-10 pairs, proximal 1 or 2 pairs anastomosing, next pair running to sinus membrane. Laminae papery, yellowish green or grayish green and pustular between veins when dried, glabrous on both surfaces. Sori orbicular, medial; indusia membranous, glabrous. Sporangia bearing light yellow glands on capsules. Spores dark yellow, irregularly thinly cristate or with small wings.

Streamsides, wet places in forests; 100-1300 m. S Fujian, Guangdong, Guangxi, S Guizhou, Hainan, S Hunan, Taiwan, SE Xizang, S Yunnan [India, Indonesia, Japan, Laos, Malaysia, Myanmar, Philippines, Sri Lanka, Thailand, Vietnam; N Australia, Pacific islands (Polynesia)].