Anictoclea grahamiana Nimmo; Tetrameles grahamiana (Nimmo) Wight; T. grahamiana var. ceylanica A. Candolle; T. rufinervis Miquel.

Trees 25-45 m tall; buttresses 2-4.5 m tall, sometimes to 6 m tall; bark gray, muricate. Petiole terete, 3-7(-20) cm; leaf blade cordate, cordate-ovate, or suborbicular, 10-26 × 9-20 cm, abaxially sparsely pubescent along veins, adaxially glabrescent, palmately 3-5-veined, margin serrate, slightly 2- or 3-lobed when young, apex acute or acuminate. Male spikes 10-20 cm; peduncles yellowish pubescent; bracts spatulate, ca. 1 mm. Female spikes 8-20 cm; peduncle pubescent. Male flowers: pedicel ca. 1 mm or very short; calyx 1.5-2 mm, lobes oblong, 3-veined, margin entire or 1- or 2-dentate, apex obtuse; filaments terete, 1-3 mm; anthers subglobose, ca. 0.5 mm. Female flowers: pedicel absent or very short (less than 1 mm); calyx slightly 4-angled, puberulous, tube fusiform, 2.5-3.5 mm, 2.5-3 mm in diam. at middle, densely brown glandular punctate outside, lobes triangular, 0.5-1 mm, 3-veined, apex acute; styles 1-2.5 mm; stigmas erect or reflexed, obovate. Capsule brown-yellow at maturity, globose-urceolate, 4-5 mm, 8-10-veined outside, sparsely brown glandular punctate. Seeds slightly compressed, less than 0.5 mm. Fl. Mar-Apr, fr. May.

Rain forests in valleys, slopes of limestone mountains; 500-700 m. S Yunnan (Jinghong, Jinping, Mengla) [Bangladesh, Bhutan, Cambodia, India (including Andaman Islands), Indonesia, Laos, Malaysia, Myanmar, Nepal, New Guinea, Sri Lanka, Thailand, Vietnam; Australia (Queensland)].