Phlomidopsis tuberosa (Linnaeus) Link; Phlomoides tuberosa (Linnaeus) Moench.

Roots enlarged, tuberous. Stems erect, 40-150 cm tall, subglabrous upward, sparsely pilose basally, sometimes purple-red. Petiole of basal leaves 4-25 cm, others 1.5-3.5 cm; basal leaf blades triangular, 5.5-19 × 5-13 cm, base deeply cordate, margin coarsely crenate, apex obtuse to acute; blade of mid stem leaves triangular-lanceolate, 5-9.5 × 2.2-6 cm, adaxially sparsely hispid to subglabrous, abaxially glabrous or sparsely bristly on veins only, base cordate, margin coarsely dentate, rarely undulate. Verticillasters ca. 3-10, widely spaced, many flowered; floral leaves short petiolate to sessile, lanceolate, reduced upward to slightly longer than verticillasters, margin dentate; bracts linear-subulate, ca. 1 cm, ciliate. Calyx tubular-campanulate, 8-10 mm, sparsely bristly near teeth otherwise glabrous outside; teeth semi-circular, 0.5-0.7 mm, apex emarginate, spine 1.8-2.5 mm. Corolla purple-red, 1.2-2 cm, tube glabrous; limb densely stellate tomentose outside, villous annulate inside; upper lip densely bearded inside, margin irregularly denticulate; lower lip ovate, ca. 6 × 5 mm; middle lobe largest, obcordate; lateral lobes ovate. Posterior filament appendages reflexed, short spurred. Nutlets apex stellate pubescent. Fl. and fr. Jul-Sep.

Wet grasslands, valleys; 1200-2100 m. Heilongjiang, Nei Mongol, Xinjiang [Kazakhstan, Kyrgyzstan, Mongolia, Russia; SW Asia, Europe].