Bark exfoliating, rough. Buds conical, slightly viscid, smooth. Leaves of short branchlets and long shoots with petiole terete or slightly compressed distally; leaf blade suborbicular, base cordate or deeply cordate. Disc parted or sinuate, persistent. Male flower: stamens 12-40; anthers long elliptic, apex apiculate. Female flower: ovary downy (or glabrous); style elongate; stigma 2- or 3-lobed. Capsule pilose, 2- or 3(or 4)-valved.