扭叶丛本藓

Anoectangium leptophyllum Broth. in Handel-Mazzetti, Symb. Sin. 4: 30. 1929. Type. China: Yunnan, Handel-Mazzetti 679 (holotype H). Anoectangium perminutum Broth., Akad. Wiss. Wien Sitzungsber., Math.-Naturwiss. Kl., Abt. 1, 133: 563. 1924. Type. China: Yunnan, Handel-Mazzetti 338 (holotype H). Anoectangium stracheyanum var. gymnostomoides (Broth. & Yas.) Wijk & Marg., Taxon 7: 288. 1958. Anoectangium gymnostomoides Broth. & Yas., Bot. Mag. (Tokyo) 19: 150. 1915. Anoectangium tortifolium Jaeg., Ber. Thätigk. St. Gallischen Naturwiss. Ges. 1869–70: 286. 1870.

Plants usually small, 5–14(–16) mm high, yellowish green, in dense tufts. Stems erect, simple or with sympodial branches; in cross section outer cortical cells small, thick-walled, central strand absent or weakly differentiated. Leaves crisped or curved-contorted when dry, erect-spreading when moist, lanceolate to linear-lanceolate, 0.8–2.0 mm x 0.15–0.30 mm, acuminate or more or less acute at the apex, always constricted above the base; margins entire above, crenulate at shoulder portion of the base because of papillose cells; costa shortly excurrent; upper and median laminal cells irregularly hexagonal to rounded, 5–12 µm x 6–10 µm, with pluripapillae; basal cells irregularly rectangular to oblong, 8–20 µm x 6–12 µm, thick-walled, smooth; abaxial superficial cells of costa above the mid-leaf linear-rectangular. Inner perichaetial leaves broad-oblong, attenuate, 0.4–1.0 mm x 0.2–0.5 mm. Setae yellowish brown, 6–10 mm long, more or less twisted; capsules erect, ovoid to shortly cylindrical, ca. 1.2 mm long; stomata present in the neck of capsule. Spores spherical, pale, brownish green, 10–13 µm in diameter.

Type. Himalayas: Boreali-occident. reg. temp., Kumaon, Strachey & Winterbottom 19 (lectotype NY); T. Thomson 207 (syntype NY).

Chinese specimens examined: ANHUI: She-xian Co., W.-F. Zheng 610 (HKAS). BEIJING: Mi-yun Co., K.-J. Guan 2b (PE). FUJIAN: Wuyi Shan (Mt.), D.-K. Li 7774 (HKAS, SHM); Xia-men City, D.-K. Li 10957 (HKAS, SHM). GUANGDONG: Le-chang Co., L. Zhang 739 (HKAS); Lian Co., L. Zhang 773 (HKAS). GUIZHOU: Gui-yang City, B.-G. Zhong 150 (HKAS). HEBEI: Xiao-wu-tai Shan (Mt.), K.-J. Guan 44 (PE). HENAN: Lin Co., D.-X. Ye 2 (HKAS); Song Co., J.-K. Liu 443 (HKAS); Xi-xia Co., Z.-Z. He 6816 (HKAS); Yu-bei Co., D.-Z. Du 1530 (HKAS). HUNAN: Da-yong Co., Zhang-jia-jie, D.-K. Li 18133 (HKAS, SHM). JIANGXI: Lu Shan (Mt.), M. Zang 211 (HKAS). JILIN: Chang-bai Shan (Mt.), C. Gao 7845 (IFSBH). SHAANXI: Hu Co., M. Wang 240 (HKAS); Tai-bai Shan (Mt.), Z.-P. Wei 6597 (HKAS). SHANXI: Ning-wu Co., C. Gao 30862 (IFSBH). SICHUAN: Ma-er-kang Co., X.-J. Li 1539 (HKAS); Omei Shan (Mt.), D.-K. Li 15571 (SHM). TAIWAN: Hsueh Shan Mo, Tungshih, Iwatsuki & Sharp 3007 (NICH); Hua-lien Co., Koponen 18335 (H); Ping-tung Co., C.-C. Chuang 1386 (MO). XIZANG: Cha-yu Co., S.-G. Wu 1097 (HKAS); Ding-ri Co., K.-Y. Lang 5303 (PE, HKAS); Lang Co., M. Zang 1898 (HKAS); Long-zi Co., M. Zang 1122 (HKAS). YUNNAN: An-ning Co., X.-J. Li 7965 (HKAS); Ge-jiu Co., X.-J. Li 4713 (HKAS); Gong-shan Co., M. Zang 1678 (HKAS); Kun-ming City, X.-J. Li 263 (HKAS); Li-jiang Co., X.-J. Li 81721 (HKAS); Yi-liang Co., X.-J. Li 4511 (HKAS). ZHEJIANG: Xi-tian-mu Shan (Mt.), M. Zang 481 (HKAS).

Habitat: often on rocks or boulders in alpine regions, near 5000 m, also at lower elevations on stone walls or in crevices; Distribution: China, Japan, Nepal, India, and Myanmar.

This species is recognized by its leaf margins constricted above the leaf base, abaxial superficial cells of costa above the mid-leaf with densely rough papillae, and linear-lanceolate leaves that are contorted when dry.

Illustrations: Pl. 107, figs. 4–6.