Andreaea rupestris ssp. fauriei Hedw., Willdenowia. 5: 24. 1968; Andreaea fauriei Besch., Ann. Sci. Nat. Bot., sér. 7, 17: 392. 1893. Andreaea rupestris Hedw. var. fauriei (Besch.) Tak., J. Hattori Bot. Lab. 11: 90. 1954.
Andreaea mamillosula Chen in Chen & Wan, Acta Phytotax. Sin. 7(2): 102. 1958. Type. China: Fujian, Wu-yi Shan, P.-C. Chen et al. 960 (holotype PE); Anhui, P.-C. Chen et al. 6872, 7577, 7575 (paratypes PE).
Andreaea rupestris ssp. fauriei is characterized by having ovate-panduriform to panduriform leaves with obtuse, incurved apices and a predominantly dioicous sexual condition. Monoicous populations were reported from Japan (Takaki 1953). The color of the plants is mostly reddish brown, at times dark brown. The papillae or mammillae of laminal cells are usually distinct.
Type. Japan, Fauriei 138 (isotype BM).
Selected Chinese specimens examined: ANHUI: Huang Shan (Mt.), see the paratypes of A. mamillosula cited above. SHAANXI: Taibei Shan (Mt.), Z.-P. Wei 5728 (WUK). ZHEJIANG: Yong-jia Co., (Yantan Shan), C. Gao 970(2) (IFSBH); Ci-xi City (Guanchuang Shan), C. Gao 687 (IFSBH).
Habitat: on rocks and boulders in mountains at high elevations; Distribution: China, Japan, and Korea.
Andreaea mamillosula described from China by Chen and Wan (1958) is identical to A. rupestris ssp. fauriei from Japan. When Bescherelle (1893) described A. fauriei from Japan, he stated that the species was differentiated from A. petrophila Ehrh. (= A. rupestris) in being dioicous and having panduriform leaves. Chen and Wan (1958) described A. mamillosula as having panduriform leaves with distinct papillae on leaf cells, but they did not mention its sexual condition. Our examination of the types and other authentic specimens of A. mamillosula shows that it is dioicous. According to our observations, papillae of laminal cells in A. rupestris can be variable; some are large and distinctive while others are so small as to be overlooked. Among the holotype and paratype specimens of A. mamillosula, most plants have distinct papillae on the leaf cells, but some are nearly smooth. Therefore, we agree with Schultze-Motel (1968) and consider A. fauriei a subspecies of A. rupestris, with A. mamillosula as a synonym. Two paratypes of A. mamillosula from Shaanxi (X.-J. Li 717, 856) are identical to A. rupestris ssp. rupestris.
Illustrations: Pl. 12, figs. 1–11.
