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Moss China | Family List | Moss China V. 1 | Dicranaceae | Campylopus

16. Campylopus umbellatus (Arnott) Paris, Index Bryol. 264. 1894; Thysanomitrion umbellatum Schwaegr. & Gaud. ex Arnott, Mém. Soc. Linn. Paris 5: 263. 1827.

Campylopus blumii (Dozy & Molk.) Bosch & Sande Lac., Bryol. Jav. 1: 81. 1858. Trichostomum blumii Dozy & Molk., Ann. Sci. Nat. Bot. sér. 3, 2: 316. 1844. Campylopus coreensis Card., Bull. Herb. Boiss. sér. 2, 7: 715. 1907. Campylopus coreensis var. amoyensis Dix. & Thér., Hong Kong Naturalist Suppl. 2: 5. 1933. Type. China: Hong Kong, Mt. Parker, Herklots B. 6 C (holotype BM). Campylopus dozyanus (C. Müll.) Jaeg., Ber. Thätigk. St. Gallischen. Naturwiss. Ges. 1870–71: 418. 1872. Dicranum dozyanum C. Müll., Syn. Musc. Frond. 1: 385. 1848. Campylopus dozyi C. Müll. ex Kindb., Enum. Bryin. Exot. 50. 1888. Campylopus ferriei Broth. in Paris, Index Bryol. Suppl. 92. 1900. Campylopus leptoneuron Broth. ex Iishiba, Nipporson Senrui Sosetsu 32 (Cat. Moss Japan). 1929. Campylopus nagasakinus Broth. in Paris, Index Bryol. Suppl. 94. 1900. Campylopus nigrescens (Mitt.) Jaeg., Ber. Thätigk. St. Gallischen. Naturwiss. Ges. 1870–71: 417. 1872, hom. illeg. Pilopogon nigrescens (Mitt.) Broth., Nat. Pflanzenfam. I(3): 336. 1901. Thysanomitrion nigrescens (Mitt.) P. de la Varde, Rev. Bryol. 49: 40. 1922. Dicranum nigrescens Mitt., J. Proc. Linn. Soc., Bot., Suppl. 1: 19. 1859. Campylopus richardii auct. non Brid. Thysanomitrion richardii auct. non (Brid.) Schwaegr. Campylopus scabripilus Warnst., Hedwigia 57: 80. 1915. Campylopus viridulus Card., Bull. Herb. Boissier sér. 2, 7: 715. 1907. Thysanomitrion sinense Broth., Nat. Pflanzenfam. ed. 2, 10: 189. 1924, nom. nud.

Plants medium-sized to large, 3–5 (–9) cm high, usually robust, yellowish green above, yellowish brown to dark brown or blackish below, somewhat shiny, in densely tomentose tufts. Stems erect or ascending, sterile ones often caudate at the apex with smaller, appressed leaves, fertile ones comose foliate, with larger, clustered and widely spreading leaves. Leaves usually appressed when dry, erect-spreading when moist, broadly lanceolate, up to 6 mm long, lamina reaching to the leaf apex, gradually narrowed above the middle, leaf base auriculate and slightly contracted; margins plane, or somewhat inflexed, mostly entire, serrulate only at the apex; costa occupying 1/3 or less of the leaf base width, long-excurrent, ending often in a short to long, serrate hyaline point, with lamellae of 2–4 cells high at back in the upper part, with both dorsal and ventral stereid bands in transverse section; upper cells obliquely oval to rhomboidal, thick-walled; basal cells subquadrate to short-rectangular, incrassate, becoming narrower at the margins; alar cells forming well marked auricles, inflated, hyaline or reddish brown. Dioicous. Perichaetial leaves elongate. Setae 4–6 mm long, aggregated, strongly flexuose to twisted when dry, cygneous when moist; capsules oblong-ovoid, ca. 1.5 mm long, brownish, apophysis distinct and mammillose; opercula long-rostrate; annuli compound, deciduous; peristome teeth linear, divided below the middle, sometimes nearly to the base, strongly papillose-striate. Calyptrae fringed at base. Spores 8–12 µm in diameter, smooth.

Type. U.S.A.: Hawaii, Gaudichaud s.n.

Chinese specimens examined: ANHUI: Huang Shan (Mt.), D.-K. Li 62475 (SHM); She- xian Co., W.-F. Zheng 1275 (MO, as Campylopus coreensis). FUJIAN: Nan-ping Co., C. Gao 25545 (MO); Sha-wu, H.-N. Fan 9282 (SYS). GUANGDONG: Lo-fu Shan (Mt.), Magill et al. 8120 (MO); Shi-xing Co., P.-J. Lin 4391(c) (IBSC). GUANGXI: Shang-si Co., C. Gao et al. 2151 (IFSBH); Xing-an Co., C. Gao & G.-C. Zhang 1895(a) (IFSBH). GUIZHOU: Jiang-kou Co., C. Gao 31251 (MO), C. Gao & J.-Y. Feng 32422 (IFSBH); Sui-yang Co., C. Gao & J.-Y. Feng 33628 (IFSBH). HAINAN: Bawangling (Mt.), C.-H. Gao 25357 (SHM); Ledong Co., Reese 17762 (MO); Li-shui Co., Redfearn 33898 (MO). HUBEI: Ba-dong Co., L.-J. Bi & J.-M. Yang 82 (IFSBH). HUNAN: Changsha City, C. Gao 97 (IFSBH); Da-yong Co., D.-K. Li 18552 (SHM); Liu-yang Co., C. Gao 653 (IFSBH). JIANGXI: Jinggong Shan (Mt.), C.-H. Gao 75341 (SHM); Lu Shan (Mt.), P.-C. Chen 60, 93 (MO). SICHUAN: Nan-chuan Co., P-C. Wu 21178 (MO, as Campylopus richardii); Omei Shan (Mt.), C.-H. Gao 161 (SHM); Tian-quan Co., C. Gao et al. 18325 (IFSBH, MO); Ya-an Co., Q. Li 750 (IFSBH), C. Gao et al. 17028 (MO). TAIWAN: Iwatsuki & Mizutani 195 (NICH); Chia-yi Co., M. -J. Lai 10082 (MO); Hwa-lien Co., C.-C. Chuang 5563 (MO); Ilan Co., C.-C. Chuang 1629 (MO); Nan-tou Co., Boufford et al. s.n., 27 June 1977 (MO); Ping-tung Co., C.-C. Chuang 1168 (MO); Tai-chung Co., M.-J. Lai 9999 (MO); Tai-pei Co., C.-C. Chuang 5442 (MO). XIZANG: Lin-zhi Co., K.-Y. Lang 304 (PE). YUNNAN: Da-li Co., Redfearn et al. 1092 (MO); Kunming City, D.-K. Li 3674 (SHM); Jing-hong Co., Redfearn 34015 (MO); Lu-feng Co., Redfearn et al. 1775 (MO); Lu-djiang (Salwin), Handel-Mazzetti s.n. (Kryptogamae exsiccatae 3080) (MO, as Thysanomitrion nigrescens); Lu-xi Co., X.-J. Li 60 (KUN); Meng-la Co., Redfearn 33889 (MO); Qiu-bei Co., D.-K. Li 2834 (SHM); Teng-chong Co., X.-J. Li 489 (KUN), 221 (MO); Wei-shan Co., Redfearn et al. 1213 (MO); Yang-bi Co., Redfearn et al. 481 (MO). ZHEJIANG: Sui-chang Co., R.-L. Hong 1659 (HSNU, IFSBH).

Habitat: on rocks or soil; Distribution: China, Korea, Japan, Sri Lanka, Indonesia, and Australia.

Illustrations: Pl. 28, figs. 1–9.


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