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16. Handelia Heimerl, Oesterr. Bot. Z. 71: 215. 1922; Tzvetkova in Schischk. & Bobrov, Fl. USSR 26: 119. 2000 (Engl. transl.); Podlech in Rech. f., Fl. Iran. Comp. IV. 158: 72. 1986; K. Bremer & Humphries in Bull. Nat. Hist. Mus. Lond. (Bot.) 23 (2): 105. 1993; K. Bremer, Asteraceae: 456. 1994.
Biennial or perennial, whitish-villose herbs. Leaves oblong, deeply 3-pinnatisect, uppermost filiform. Capitula homogamous, discoid, numerous, in congested corymbs. Involucre hemispherical, phyllaries ± equal. Receptacle convex-hemispherical, chaffy in upper part, paleae linear-lanceolate, obtuse, membranous. Ray-florets absent. Disc-florets bisexual, with ± cylindrical, upwardly gradually dilated, 5-toothed, campanulate, sparsely punctate-glandular corolla tube. Cypselas lengthwise prominently 5-ribbed, ± truncate at the apices. Pappus short coronate.
A monotypic Central Asian genus, recorded for the first time from Pakistan.
Lower Taxon
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