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Pakistan | Family List | Pakistan V. 203 | Salicaceae

1. Salix Linnaeus, Sp. Pl. 1015. 1753; Gen. Pl. ed. 5: 447. 1754; Andersson, Salicaceae in DC., Prodr. 16(2): 190. 1868; Boiss., Fl. Or. 4: 1181. 1879; Hook.f., F. Brit. Ind. 5: 626. 1888; R. Parker, For. Fl. Punj. 3rd. ed. 498. 1956; A. K. Skvortsov in Rech. f., Fl. Iran. 65: 12. 1969; F. Zhenfu, Z. Shidong & A. K. Skvortsov in Wu Zheng-yi & P.H.Raven, Fl. China 4: 162. 1999.

Trees, shrubs or subshrubs. Buds with single outer scale. Leaves alternate, stipules often present, petiole short or absent; lamina linear lanceolate, oblong or obovate. Catkins appearing before, with or after the leaves; erect, spreading or rarely pendulous, sessile or pedunculate. Flower bracteate, bract 1. Glands 1 or 2 (or more) small. Stamens usually 2 (sometimes up to 12). Filaments free or united. Ovary unilocular, mostly stipitate; style and stigmas variable. Fruit 2-valued; valves mostly recurving after dehiscence. Seeds small, comose.

A genus with c. 520 species; represented in Pakistan by c. 26 species.

As the male and female plants exist separately, and, in some cases considerable hyberidization between different taxa is also involved, hence taxonomically Salix is a difficult genus. Considerable morphological variation is met with in each species.

Note: Salix nuristanica A.K.Skvortsov could not be keyed due to lack of male plants.


Key based on female plants

1 Capsule glabrous.   (2)
+ Capsule hairy.   (16)
       
2 (1) Dwarf shrubs of alpine areas; leaves generally not exceeding 2.5 cm.   (3)
+ Small or large shrubs or trees.   (4)
       
3 (2) Rachis of the female catkin hairy; catkin 10-45 mm long; leaf margin crenate or serrate.   18 S. flabellaris
+ Rachis of the female catkin glabrous; cakin 7.5-12.7 mm long; leaf margin entire.   19 S. lindleyana
       
4 (2) Branches long drooping, tree with a ‘weeping’ habit.   (5)
+ Branches erect spreading, not drooping.   (6)
       
5 (4) Leaves in the catkin producing region much smaller. Nectary one, longer than the stipe.   13 S. babylonica
+ Leaves in the catkin producing region ± of the same size. Nectaries 1-2 per flower, equalling the fruit stipe (0.2-0.8 mm).   12 S. alba
       
6 (4) Bract, ovary and fruit reddish violet.   11 S. karelinii
+ Bract, ovary and fruit not as above.   (7)
       
7 (6) Branches mostly covered with white bloom, stipe of the ovary c. 1 mm long.   5 S. capusii
+ Branches not covered by the white bloom.   (8)
       
8 (7) Fruiting catkin c. 1 cm long.   26 S. nuristanica
+ Fruiting catkin >1 cm long.   (9)
       
9 (8) Lamina ovate or elliptic-oblong, tip obtuse, sometimes acute; stigma subsessile.   9a subsp. denticulata
+ Lamina not as above.   (10)
       
10 (9) Leaf elliptic or oblong-obovate, tip shortly acuminate, usually oblique.   23 S. fedtschenkoi
+ Leaf lanceolate to linear lanceolate or ovate lanceolate or elliptic or broadly elliptic, tip not oblique.   (11)
       
11 (10) Fruiting catkin 5 cm or more in length.   (12)
+ Fruiting catkin mostly less than 5 cm in length.   (14)
       
12 (11) Fruiting catkin 10-18 cm in length; stipules up to 1.3 cm long.   16 S. dolichostachya
+ Fruiting catkin 10 cm or less in length stipules < 1 cm long.   (13)
       
13 (12) Fruiting catkin lax; fruit stipe 0.5-3 mm, capsule 3.5-5 mm long.   2 S. tetrasperma
+ Fruiting catkin not lax; fruit stipe 0.1-0.8 mm, capsule 4-6 mm long.   6 S. pycnostachya
       
14 (11) Lamina broadly elliptic, shortly acuminate.   14 S. excelsa
+ Lamina not as above.   (15)
       
15 (14) Fruit stipe 0.1-0.8 mm, capsule 4-6 mm.   6 S. pycnostachya
+ Fruit stipe 0.4-1.8 (-2.5) mm, capsule 3-5.5 mm.   1 S. acmophylla
       
16 (1) Lamina about half as broad as long; margin sinuate-erose. Capsule 7-9 mm.   22 S. aegyptiaca
+ Lamina not as above. Capsule <= 7 mm.   (17)
       
17 (16) Lamina elliptic to subrhomboidal, rarely ovate, narrowed at the base, tip shortly acuminate (capsule stipe 1-3 mm).   15 S. iliensis
+ Lamina not as above.   (18)
       
18 (17) Lamina elliptic, oblong or ovate, 2.5-6.5 x 1.2-2.4 cm, tip obtuse or acute.   9b subsp. hazarica
+ Lamina not as above.   (19)
       
19 (18) Branches mostly covered by white bloom.   5 S. capusii
+ Branches not covered by white bloom.   (20)
       
20 (19) Leaf silky on the lower side.   (21)
+ Leaf not silky on the lower side.   (24)
       
21 (20) Catkin 5-8 cm long.   17 S. wallichiana
+ Catkin 1.5-5 cm long.   (22)
       
22 (21) Leaf (linear-) lanceolate, obovate, villous. Capsule 5-7 mm long.   24 S. schugnanica
+ Leaf linear, lanceolate or oblong to ovate-oblong, not villous. Capsule 4-6 mm long.   (23)
       
23 (22) Leaf linear, lanceolate, (5)-12-20 x 0.5-2-(4) cm, lateral veins 25-30 pairs.   20 S. viminalis
+ Leaf broadly lanceolate, oblong to ovate-oblong, 4.5-14 x 1-2.5 cm, lateral veins 16-18-(20) pairs.   21 S. turanica
       
24 (20) Catkin 2 cm or less long.   (25)
+ Catkin 3 cm or more long.   (28)
       
25 (24) Lamina generally more than 1 cm broad.   (27)
+ Lamina 9 mm or less broad.   (26)
       
26 (25) Branches pubescent or silky, rarely glabrate. Leaf exstipute, hairy on the sides, becoming glabrescent, 2-4 mm broad.   3 S. wilhelmsiana
+ Branches glabrous. Leaves stipulate, glabrous or the lower side in young leaves scantily silky, 3-7-(9) mm broad.   4 S. koeieana
       
27 (25) Lamina oblong, elliptic or lanceolate. Stigma entire or 2-lobed. Gland 1.   10 S. caesia
+ Lamina elliptic, ovate or ovate to suborbicular. Stigma 4-lobed. Glands 2.   25 S. sclerophylla
       
28 (24) Lamina 1 cm or less broad.   (29)
+ Lamina generally broader than 1 cm.   (30)
       
29 (28) Ovary and capsule sessile, stigma entire or bilobed.   3 S. wilhelmsiana
+ Ovary and capsule stipitate, stigmas 4 as upright lobes.   7 S. blakii
       
30 (28) Stipules minute or semicordate, coarsely toothed. Stigmas 2-4, longer than the style. Capsule stipe 0.7-2 mm long; fruit 5-9 mm long.   17 S. wallichiana
+ Stipules linear, lanceolate. Stigmas 2-4, shorter or equal to the style. Capsule stipe 0, 1-10 mm long; fruit 4-6 mm long.   (31)
       
31 (30) Stipules persistent.   8 S. sericocarpa
+ Stipules not persistent.   6 S. pycnostachya

List of Keys

  • List of lower taxa


     

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