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11. Tanacetum Linnaeus, Sp. Pl. 843. 1753 pro parte; Gen. Pl. ed. 5. 366. 1754 emend. Tzvelev in Schischk. & Bobrov, Fl. USSR 26: 303. 2000 (Engl. transl.); Podlech in Rech. f., Fl. Iran. Comp. IV. 158: 1986- pro parte; K. Bremer & Humphries in Bull. Nat. Hist. Mus. Lond. (Bot.) 23(2): 100. 1993; K. Bremer, Asteraceae: 465. 1994; Grierson & Springate in Grierson, D. G. Long & Springate, Fl. Bhutan 2 (3): 1549. 2001.
Spathipappus Tzvelev in Schischk. & Bobrov, Fl. USSR 26: 875. 1961; Podlech in Rech. f., Fl. Iran. Comp. IV. 158: 148. 1986.
Perennial or rarely annual herbs, usually aromatic, often rhizomatous and basally woody shrublets with erect or ascending, leafy and branched, sometimes subscapiform shoots. Leaves alternate, pinnatifid or 1-3-pinnatisect, punctate, rarely entire, serrate. Capitula heterogamous, radiate, disciform, or discoid, solitary or corymbose. Involucre hemispherical or shallowly to broadly campanulate, phyllaries 3-4-seriate, imbricate, with scarious or sometimes with dark brown margins. Receptacle flat, hemispherical or convex, epaleate. Ray-florets, when present, female, fertile or sterile, with 3-lobed, white, yellow or pink ligules, in discoid forms tubular, female or hermaphrodite. Disc-florets tubular, 5-lobed, yellow. Anthers minutely sagitatte at the bases, apically ovate-appendaged. Cypselas homomorphic, flat, usually oblong, 5-12-ribbed, glabrous or with sessile glands, without myxogenic cells or secretory canals. Pappus a corona or usually unevenly toothed, scaly or lobed, sometimes adaxially well-developed.
A fairly large genus comprising c. 152 species, distributed mainly in Europe and temperate Asia, N Africa, N America and in the southern hemisphere by introduction (Humphries, 1993). Represented in Pakistan by 12 species, mainly confined to mountainous areas in North and North West.
Co-author: Ronaq Khan, Natural History Museum, London.
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| 1 |
Capitula discoid, with all florets tubular. |
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(11) |
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Capitula radiate, with marginal florets ligulate. |
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(2) |
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| 2 (1) |
Ligules white. |
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(3) |
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Ligules pale pink to reddish-pink. |
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9 Tanacetum coccineum |
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| 3 (2) |
Capitula solitary, terminal. |
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(4) |
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Capitula few (3) to many (30), in lax corymbs. |
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2 Tanacetum parthenium |
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| 4 (3) |
Ligulate florets sterile, with ovaries jointed with the corolla bases. |
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(10) |
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Ligulate florets fertile, with the ovaries not jointed with the corolla bases. |
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(5) |
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| 5 (4) |
Pappus of obtuse, membranous scales. |
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3 Tanacetum pakistanicum |
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Pappus of a short corona or deltate rigid teeth. |
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(6) |
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| 6 (5) |
Leaves divided less than halfway to midrib; capitulum broader than long. |
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4 Tanacetum emodi |
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Leaves divided more than halfway to midrib; Capitula longer than broad. |
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(7) |
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| 7 (6) |
Leaves clustered around the inflorescence. |
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5 Tanacetum roylei |
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Leaves not clustered around the inflorescence, mostly in the middle or lower half of the stem. |
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(8) |
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| 8 (7) |
Leaves densely silvery-gray pubescent on both sides, uppermost leaves dissected. Involucre 12 – 18 mm in diameter. |
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8 Tanacetum cinerariifolium |
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Leaves glabrescent to glabrous, uppermost leaves linear, entire. Involucre less than 10 mm in diameter. |
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(9) |
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| 9 (8) |
Leaves pinnatisect. Pappus shortly coronate, c. 0.25 (- 0.5) mm long. |
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6 Tanacetum stoliczkae |
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Leaves pinnatifid. Pappus of irregular teeth and linear, free appendages. |
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7 Tanacetum baltistanicum |
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| 10 (4) |
Leaves 20 – 50 mm long, 2-3-pinnatisect. Phyllaries with dark scarious margins. Pappus more than 1.5 mm long. |
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11 Tanacetum griffithii |
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Leaves 5 – 10 mm long, unipinnatisect. Phyllaries with pale puberulous margins. Pappus less than 1 mm long. |
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12 Tanacetum chitralense |
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| 11 (1) |
Leaves more than 4 cm long, with long petioles. |
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1 Tanacetum falconeri |
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Leaves less than 3 cm long, with inconspicuous or short petioles. |
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10 Tanacetum artemisioides |
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List of lower taxa
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