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Pakistan | Family List | Amaranthaceae | Amaranthus

Amaranthus retroflexus Linn., Sp. Pl. 991. 1753. Boiss., Fl. Or. 4:989. 1879; Stewart, Ann. Cat. Vase. Pl. W. Pakistan: 231. 1972; Aellen in Rech. f., Fl. Iranica Lf. 91: 3. 1972.

Annual herb, erect or with ascending branches, (6-) 15-80 (-100) cm, simple or branched (especially from the base to about the middle of the stem). Stem stout, sub-terete to angled, densely furnished with multicellular hairs. Leaves furnished with multicellular hairs along the lower surface of the primary venation and often the lower margins, long-petiolate (petioles up to c. 6 cm, in robust plants not rarely equalling the lamina), lamina ovate to rhomboid or oblong-ovate, (1-) 5-11 x (0.6-) 3-6 cm, obtuse to subacute at the mucronulate tip, shortly cuneate or attenuate into the petiole. Flowers in greenish or rarely somewhat pink-suffused, stout, axillary and terminal spikes, which are usually shortly branched to give a lobed appearance, more rarely with longer branches, the terminal inflorescence paniculate, very variable in size, male and female flowers intermixed, the latter generally much more plentiful except sometimes at the tip of the spikes. Bracts and bracteoles lanceolate-subulate, pale-membranous with a prominent green midrib excurrent into a stiff, colourless arista, longer bracteoles subequalling to twice as long as the perianth. Perianth segments 5, those of the male flowers 1.75-2.25 mm, lanceolate-oblong, blunt to subacute, those of the female flowers 2-3 mm, narrowly oblong-spathulate to spathulate, obtuse or emarginate, ± green-vittate along the midrib, which ceases below the apex or is excurrent in a short mucro. Stigmas 2-3, patent-flexuose or erect, c. 1 mm. Capsule subglobose, c. 2 mm, usually shorter than the perianth, circumscissile, with an indistinct neck, rugose below the lid. Seed black and shining, compressed, c. 1 mm, almost smooth centrally, faintly reticulate around the margins.

Type: Cultivated material from Uppsala Botanic Garden, Linnean specimen 1117/22 (LINN, holotype!).

Distribution: A native of N. America south to N. Mexico; introduced into the Old World as a weed, but in more temperate regions than many of its allies, occurring in S. and C. Europe, Mediterranean N. Africa and temperate Asia from Cyprus and Turkey to Iran, Siberia, Middle Asia, Mongolia, China and Japan. Also adventive in Australia and S. America (Bolivia etc.) and probably elsewhere. Generally occurs as a weed of cultivation, occurring as such in the cooler parts of Pakistan, ascending to 1820 m or more.


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