Brothera J. K. A. Müller, Gen. Musc. Frond. 258. 1901.
[for Viktor Ferdinand Brotherus, Finnish bryologist]
Plants small, 3--6 mm high, in dense, yellowish to grayish green mats. Stems radiculose at base. Leaves erecto-patent when dry, lanceolate, gradually contracted to a short, canaliculate apex; costa filling 2/3 of the leaf base, not sharply delimited from the lamina, excurrent, in transverse section with lax abaxial and adaxial hyalocysts and a median band of stereids; alar cells weakly differentiated, hyaline; basal laminal cells hyaline, of 8--10 rows, 2-stratose towards the costa, rectangular; distal laminal cells rectangular. Sexual condition dioicous. Seta 5--6 mm, yellowish, sinuose and twisted sinistrorse in the distal portion. Capsule erect, yellowish green, 0.5 ´ 1.2 mm, smooth, contracted towards the peristome; annulus present; operculum rostrate, half the length of the urn; peristome teeth 16, entire or rarely perforated, yellowish, striate at base and papillose at tips, 210--240 µm; spores yellowish green, smooth, 10--13 µm; exothecial walls incrassate, variable, without stomata. Calyptra cucullate, fringed at base.
Species 1 (1 in the flora): a disjunctive genus of se North America, Central Ameica, Asia, Africa.
Müller, P. & Frahm, J.-P. 1987. A review of the Paraleucobryoideae (Dicranaceae). Nova Hedwigia 45: 283-314.