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BFNA | Family List | BFNA Vol. 1 | Dicranaceae | Campylopus

Campylopus introflexus (Hedwig) Bridel, Mant.Musc. 72. 1819.

  • Dicranum introflexum Hedwig

    Plants 0.5--5 cm, in dense mats, yellowish to olive green, tomentum present or almost absent. Leaves 4--6 mm, erect patent when wet, appressed when dry, lanceolate, straight, with entire margins; alar cells lacking or formed by thin-walled, hyaline to reddish, inflated cells; basal laminal cells hyaline rectangular, thin-walled, extending higher at margins and forming a V-shaped area; distal laminal cells incrassate, shortly rectangular to oblique, chlorophyllose; costa filling 1/2--3/4 of leaf width, excurrent in a hyaline hair tip, which is conspicuously 90° reflexed, in transverse section showing adaxial hyalocysts and abaxial stereids, shortly lamellose at back with ribs 1--2 cells high. Specialized asexual reproduction occasionally by deciduous stem tips. Seta 7--12 mm, yellowish brown to brownish in age, often several sporophytes from the same plant, curved or sinuose. Capsule brown, 1.5 mm, slightly asymmetric and curved when empty. Calyptra ciliate at base. Spores 12--14 µm.

    Soil along trails, base of trees, flat roofs of buildings, peat in bogs, sand; 0--200 m; B.C.; Calif, Oreg., Wash.; Europe; South America (Chile, Argentina, se Brazil); s Africa, Australia, Pacific Islands (New Caledonia, New Zealand), Subantarctic Islands.

    This species occurs in masses in sand dunes along the west coast of North America and throughout the Southern Hemisphere. The species was introduced in Britain in 1942, since the beginning of the 1970's has been aggressively spreading through Europe and is now ranging from Iceland to Spain and from Ireland to Poland. The first record in North America dates from August, 1975, and was made on a gravel roof of a building of Humboldt University, Arcata, California. The species is undoubtedly introduced in North America and is spreading as rapidly as in Europe. The name C. introflexus was used previously for C. pilifer, thus all old references for C. introflexus in North America have to be referred to that species. Also specimens of C. surinamensis and C. oerstedianus from North America were named as C. introflexus. Campylopus introflexus is easily recognized by the reflexed hair points. Female plants have terminal perichaetial buds. Problems may rarely arise with forms from shaded habitats, in which the hairpoints are lacking or so short that they are not reflexed.


     

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