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BFNA | Family List | BFNA Vol. 1 | Pottiaceae | Tortula

Tortula modica R. H. Zander, Bull. Buffalo Soc. Nat. Sci. 32: 226. 1993.
[new name]

  • Gymnostomum intermedium Turner
  • Pottia intermedia (Turner) Fürnrohr
  • Pottia truncata var. major (F. Weber & D. Mohr) Bruch & Schimper

    Leaves obovate to spatulate, apex broadly acute or occasionally rounded, short-awned, margins weakly recurved proximally or occasionally plane, weakly bordered distally with 2--4 rows of slightly thicker-walled cells; costa excurrent, lacking an adaxial pad of cells, distally narrow, 2--3 cells across the convex adaxial surface; distal laminal cells irregularly hexagonal or sometimes rectangular, width 20--25(--29) µm wide, 1(--2):1, smooth or 1-papillose (best seen in section). Sexual condition autoicous. Sporophytes exerted. Seta 0.4--0.6 cm. Capsules stegocarpic, not systylius, short-cylindric, erect and nearly straight, urn 1--1.5 mm; peristome absent or rudimentary; operculum ca. 0.5 mm. Spores 30--35 µm, spheric, densely papillose.

    Capsules mature fall--spring. Calcareous soil, old fields, pastures, rocky areas; low and moderate elevations; Ont. (Bruce, Durham, Grey, Victoria counties); N.Y., Pa.; Europe; Asia; n Africa.

    A report from Mexico was excluded by R. H. Zander (1993). The distribution and characters of this uncommon species are discussed by H. A. Crum and L. E. Anderson (1981), P. M. Eckel (1987), C. Williams (1966). Essentially, Tortula modica is a high polyploid (n = 52) taxon distinguished from T. truncata by the larger habit, leaf margins usually recurved proximally, capsules slightly longer, short-cylindric, not tapering to the base, and peristome sometimes present though rudimentary.


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