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Tortula guepinii (Bruch & Schimper) Brotherus, Natürl. Pflanzenf. 1(3): 430. 1902.
Desmatodon guepinii Bruch & Schimper
Leaves ovate to short-elliptic, apex broadly acute, awned, margins recurved in mid 4/5 of leaf, usually bordered distally with 2--4 rows of smooth or less papillose, somewhat thicker-walled cells; costa excurrent in the awn, lacking an adaxial pad of cells, distally narrow, 2--4 cells across the convex adaxial surface; distal laminal cells hexagonal, 16--19 µm wide, 1:1, papillose with 4--6 1- to 2-fid papillae. Sexual condition autoicous. Sporophytes exerted. Seta 0.7--0.8 cm. Capsules stegocarpic, not systylius, cylindric, erect and nearly straight, urn 1.2--1.4 mm; peristome ca. 500 µm, of 16 teeth, weakly twisted, divided to below the middle, basal membrane to 75 µm high; operculum ca. 0.8 mm. Spores 13--18 µm, spheric, papillose.
Capsules mature spring. On soil; low elevations; Ariz., Calif., Colo., Nev., N.Mex., Tex., Wyo.; Mexico (Baja California); sw Europe.
Tortula guepinii resembles T. hoppeana. of montane and subarctic distribution in North America and elsewhere, but it is smaller, with leaves less than 2 mm, and cells of the leaf margin are distinctly less papillose at the apex. Tortula plinthobia of the southeastern and southwestern U.S.A. has distal laminal cells only 10--13 µm wide.
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