Perennials. Stems pilosulous to villous and/or stipitate-glandular. Leaves: petioles 3–8(–12+) cm, sometimes raggedly winged; blades 4–20(–40+) × 2–12(–25+) cm, larger usually deeply pinnately lobed, lobes 5–7. Cypselae 3–4 mm, 3-angled or -ribbed. 2n = 30.

Flowering May–Aug(–Oct). Damp, shaded sites, calcareous soils; 100–1000 m; Ont.; Ala., Ark., Conn., D.C., Ga., Ill., Ind., Iowa, Kans., Ky., Mich., Minn., Mo., N.Y., N.C., Ohio, Okla., Pa., Tenn., Vt., Va., W.Va., Wis.