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FNA | Family List | FNA Vol. 19, 20 and 21 | Asteraceae | Cirsium

59. Cirsium fontinale (Greene) Jepson, Fl. W. Calif. 505. 1901.

Fountain thistle

Cnicus fontinalis Greene, Bull. Calif. Acad. Sci. 2: 151. 1886

Monocarpic perennials, 50–220 cm; woody tap-rooted caudices with numerous coarse, fibrous lateral roots, sometimes forming new rosettes from root sprouts. Stems 1–several, erect, loosely arachnoid-tomentose, glandular-pilose; branches several to many, spreading. Leaves: blades oblanceolate to oblong or elliptic, 20–70 × 5–16 cm, strongly undulate, shallowly to deeply pinnatifid, lobes coarsely toothed or with triangular secondary lobes, main spines 3–15 mm, abaxial faces more densely tomentose, adaxial densely glandular with short, multicellular trichomes and ± tomentose with fine, arachnoid, non-septate trichomes; basal often present at flowering, petiolate; principal cauline well distributed, becoming sessile, gradually reduced distally, bases auriculate-decurrent with broad, spiny-margined wings 3 cm or less; distal cauline leaves progressively reduced, ± bractlike. Heads few–many, ± nodding, in paniculiform or corymbiform arrays, bracts leafy or much reduced. Peduncles 0–7 cm. Involucres (green to purple), hemispheric or campanulate, 1.5–3 × 2–5 cm, glabrous or puberulent. Phyllaries in 6–10 series, imbricate, ovate to lanceolate, abaxial faces without glutinous ridge; outer and mid bases appressed, short, bodies entire to erose or ciliolate (rarely outer spine-margined), apices spreading to recurved or reflexed, long, dilated, adaxially puberulent, spines 1–6 mm; apices of inner erect or reflexed, flattened or tipped with short spines. Corollas white to pinkish or lavender, 14.5–22 mm, tubes 5–8 mm, throats 4.5–9 mm, lobes 3–8 mm; style tips 2.5–4.5 mm. Cypselae brownish, 3.4–5 mm, apical collars tan; pappi 12–15 mm.

Varieties 3 (3 in the flora): c California.

1 Cypselae 4.1–5 mm; phyllary spines 1–2 mm   59b Cirsium fontinale var. fontinale
+ Cypselae 3.4–4.1 mm; phyllary spines 2–6 mm   (2)
2 (1) Longer spines of cauline leaves 10–18 mm; most phyllaries without marginal spines   59a Cirsium fontinale var. campylon
+ Longer spines of cauline leaves 4–7 mm; many outer phyllaries with marginal spines   59c Cirsium fontinale var. obispoense


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