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FNA | Family List | FNA Vol. 19, 20 and 21 | Asteraceae | Malacothrix

6. Malacothrix foliosa A. Gray in A. Gray et al., Syn. Fl. N. Amer. ed. 2. 1: 455. 1886.

Leafy desertdandelion

Annuals, 4–45 cm. Stems 1, erect, simple or branched proximally and/or distally or 2–10, decumbent to ascending, and ± branched distally, glabrous or sparsely arachnose (sometimes only in leaf axils). Cauline leaves: proximal oblanceolate to narrowly obovate, usually pinnately lobed (lobes usually unequal, apices acute or obtuse), sometimes ± fleshy, ultimate margins entire or dentate, faces glabrous (proximalmost cauline usually more deeply divided); distal seldom notably reduced (sometimes pinnately lobed). Calyculi 0 (i.e., outer phyllaries intergrading with inner), or of 5–12+, oblong or ovate to lanceolate or linear bractlets, hyaline margins 0.1–0.3 mm wide. Involucres ± campanulate, 5–12 × 2–7 mm. Phyllaries 12–22(–40+, without calyculi) in 2–3(–5+) series, (usually red-tinged) lanceolate to linear, hyaline margins 0.05–0.2 mm wide, faces glabrous. Receptacles not bristly. Florets 10–123; corollas light to medium yellow, 5–17 mm; outer ligules exserted 1–10 mm. Cypselae ± cylindro-fusiform or ± prismatic, 0.9–1.7 mm, ribs extending to apices, ± equal or 5 more prominent than others; persistent pappi usually 0, rarely of 1–2 bristles. Pollen 70–100% 3-porate. 2n = 14.

Subspecies 4 (4 in the flora): Channel Islands, California.


1 Outer ligules usually exserted 1–4 mm   (2)
+ Outer ligules exserted 5–15 mm   (3)
       
2 (1) Calyculi 0 or bractlets intergrading with phyllaries; cypselae ± prismatic (5-angled), 1.3–1.6 mm, 5 ribs more prominent than others; Anacapa Island, California   6a Malacothrix foliosa subsp. crispifolia
+ Calyculi of 8–12+, oblong to linear bractlets; cypselae ± cylindro-fusiform, 0.9–1.5 mm, ribs ± equal; San Nicolas Island, California   6b Malacothrix foliosa subsp. polycephala
       
3 (1) Stems 1, usually erect; distal cauline leaves usually pinnately lobed (near bases, lobes 1–2 pairs, narrow); cypselae ± cylindro-fusiform, 0.9–1.5 mm, ribs ± equal; San Clemente Island, California   6c Malacothrix foliosa subsp. foliosa
+ Stems (1–)3–5(–10), erect or decumbent to ascending; distal cauline leaves usually pinnately lobed (from bases to near apices); cypselae ± prismatic (5-angled) 1.3–1.7 mm, 5 ribs more prominent than others; Santa Barbara Island, California   6d Malacothrix foliosa subsp. philbrickii


 

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