Coelophragmus umbrosus (B. L. Robinson) O. E. Schulz; Dryopetalon runcinatum var. laxiflorum Rollins; Sisymbrium umbrosum B. L. Robinson

Annuals or biennials; usually sparsely to densely hirsute proximally, rarely glabrous throughout. Stems erect, unbranched or branched basally, branched distally, 2-8 dm. Basal leaves: petiole (1-)2-6(-8) cm; blade spatulate to obovate in outline, lyrate, pinnatifid, or bipinnately lobed, (2-)4-20(-25) cm × 30-80 mm, surfaces glabrous or hirsute. Cauline leaves shortly petiolate or sessile; blade base not auriculate, similar to basal, smaller distally. Racemes lax or dense. Fruiting pedicels divaricate-ascending, 5-15(-20) mm, glabrous or pubescent. Flowers: sepals (2.5-)3.5-4.5(-5) × 1.5-2 mm; petals white to purplish, spatulate, 6-9(-11) × 2.5-3.5(-4.5) mm, claw 2.5-4(-5) mm, margins pinnatifid, 5-7(-11)-lobed, papillate basally; filaments papillate basally, median pairs 3-5 mm, lateral pair 2.5-4 mm; anthers oblong, 0.7-1 mm. Fruits linear, (straight or arcuate), 2-6 cm × 0.5-1.2 mm; ovules 60-110 per ovary; style 0.1-0.7(-1) mm. Seeds 0.6-0.8 × 0.4-0.6 mm, not winged. 2n = 24.

Flowering Jan-May. Ledges, shade of boulders and cliffs, foothills, canyons, scrub woodlands, streambeds, rocky basalt, crevices; 800-1900 m; Ariz., N.Mex., Tex.; Mexico (Chihuahua, Sinaloa, Sonora).

Dryopetalon runcinatum is easily distinguished from other mustards in the flora area by having divided petals. In the United States, it is restricted to Cochise, Pima, Pinal, and Santa Cruz counties in Arizona, Catron, Dona Ana, Hidalgo, and Luna counties in New Mexico, and El Paso County in Texas.