32. Rubus setosus Bigelow, Fl. Boston. ed. 2. 198. 1824.
Bristly blackberry, ronce sétuleuse
Rubus apparatus L. H. Bailey; R. beatus L. H. Bailey; R. benneri L. H. Bailey; R. bicknellii L. H. Bailey; R. boottianus L. H. Bailey; R. condignus L. H. Bailey; R. deaneanus L. H. Bailey; R. discretus L. H. Bailey; R. dissensus L. H. Bailey; R. dissimilis L. H. Bailey; R. electus L. H. Bailey; R. exter L. H. Bailey; R. fulleri L. H. Bailey; R. groutianus Blanchard; R. gulosus L. H. Bailey; R. hispidoides L. H. Bailey; R. hispidus Linnaeus var. suberectus Peck; R. jejunus L. H. Bailey; R. junceus Blanchard; R. junior L. H. Bailey; R. lawrencei L. H. Bailey; R. mediocris L. H. Bailey; R. navus L. H. Bailey; R. nocivus L. H. Bailey; R. notatus L. H. Bailey; R. ortivus (L. H. Bailey) L. H. Bailey; R. perinvisus L. H. Bailey; R. racemiger L. H. Bailey; R. regionalis (L. H. Bailey) L. H. Bailey; R. reravus L. H. Bailey; R. rotundior (L. H. Bailey) L. H. Bailey; R. schneideri L. H. Bailey; R. semisetosus Blanchard; R. spectatus L. H. Bailey; R. stipulatus L. H. Bailey; R. uniformis L. H. Bailey; R. univocus L. H. Bailey; R. vermontanus Blanchard; R. viridifrons L. H. Bailey; R. wheeleri (L. H. Bailey) L. H. Bailey
Shrubs, 2–10(–15) dm, armed. Stems ˂biennial˃, erect to arching, rarely creeping, ˂not node- or tip-rooting˃, sparsely to moderately hairy, sparsely to densely long-, rarely short-stipitate-glandular, ˂glands rounded to flattened˃, not pruinose; prickles absent or sparsely to moderately dense, erect to retrorse, weak, slender, 1–3(–5) mm, narrow-based; ˂bristles sparse to dense, erect to retrorse, green to reddish, narrow, semirigid, sometimes flexible, not gland-tipped˃. Leaves deciduous, ternate or palmately compound, ˂sometimes ± lustrous˃; stipules filiform to lanceolate, 5–20(–38) mm; leaflets (3–)5, terminal obovate or elliptic to rhombic, 4.5–11.5 × 2–7 cm, base cuneate to narrowly rounded, unlobed, margins moderately, coarsely serrate to doubly serrate or serrate-dentate, apex acute or acuminate to short-attenuate, abaxial surfaces unarmed or with bristles on midvein, sparsely to moderately hairy, eglandular or sparsely to moderately sessile- to long-stipitate-glandular along larger veins. Inflorescences terminal ˂on short shoots, sometimes appearing axillary˃, 5–15(–20)-flowered, racemiform, cymiform, or thyrsiform. Pedicels prickles or bristles sparse to dense, erect to retrorse, moderately to densely hairy, moderately to densely stipitate-glandular. Flowers bisexual; petals white, obovate to oblanceolate, 6–13 mm; filaments filiform; ovaries glabrous. Fruits black, globose, 0.7–1.5 cm; drupelets 5–25, strongly coherent, separating with torus attached. 2n = 14, 21, 28, 35.
Flowering Jun–Jul(–Aug). Open woodlands, savannas, prairies, meadows, disturbed areas, dry to wet soil; 0–1000 m; N.B., Nfld. and Labr. (Nfld.), N.S., Ont., P.E.I., Que.; Conn., Del., D.C., Ill., Ind., Iowa, Maine, Md., Mass., Mich., Minn., N.H., N.J., N.Y., Ohio, Pa., R.I., Vt., Va., W.Va., Wis.
Rubus setosus is identified by erect to arching, rarely creeping, not tip-rooting, terete primocanes, always setose but variable in density, and deciduous primocane leaves that are often palmately compound with five leaflets. Cultivated plants sometimes alternate from erect to creeping (or vice-versa) in successive years, depending on changing soil conditions.
The following nothospecies names are based on putative hybrids involving Rubus setosus and: R. allegheniensis (R. ×abbrevians Blanchard, R. ×aculiferus Fernald, R. ×adenocaulis Fernald, R. ×angustifoliatus L. H. Bailey, R. ×ascendens Blanchard, R. ×atwoodii L. H. Bailey, R. ×clausenii L. H. Bailey, R. ×flavinanus Blanchard, R. ×frondisentis Blanchard, R. ×perspicuus L. H. Bailey, R. ×ravus L. H. Bailey); R. canadensis (R. ×miscix L. H. Bailey [based on R. ×peculiaris Blanchard (not R. peculiaris Sampaio)]); R. flagellaris (R. ×alter L. H. Bailey, R. ×arcuans Fernald & H. St. John, R. ×biformispinus Blanchard, R. ×bigelovianus L. H. Bailey, R. ×bracteolifer Fernald, R. ×jacens Blanchard var. specialis L. H. Bailey, R. ×laevior (L. H. Bailey) Fernald [based on R. permixtus var. laevior L. H. Bailey], R. ×multiformis Blanchard, R. ×setospinosus L. H. Bailey, R. ×severus Brainerd ex Fernald); R. hispidus (R. ×adjacens Fernald, R. ×grandidens L. H. Bailey, R. ×harmonicus L. H. Bailey, R. ×jacens Blanchard, R. ×parlinii L. H. Bailey, R. ×spiculosus Fernald, R. ×tardatus Blanchard, R. ×tholiformis Fernald, R. ×trifrons Blanchard, R. ×zaplutus L. H. Bailey); R. pensilvanicus (R. ×wisconsinensis L. H. Bailey).