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FNA | Family List | FNA Vol. 9 | Rosaceae | Horkelia

12b. Horkelia californica Chamisso & Schlechtendal var. elata (Greene) Ertter & Reveal, Novon. 17: 318. 2007.

Tall horkelia

Potentilla californica (Chamisso & Schlechtendal) Greene var. elata Greene, Fl. Francisc. 1: 66. 1891, based on P. elata Greene, Pittonia 1: 100. 1887, not Salisbury 1796; Horkelia californica subsp. dissita (Crum) Ertter; H. elata (Greene) Rydberg; P. elata var. dissita Crum

Basal leaves (5–)10–25 cm; sheathing bases usually glabrous, sometimes sparsely strigose; lateral leaflets (5–)7–9 per side, ovate to round, 5–30 mm, cleft 1/2–3/4+ to midrib, ultimate teeth 10–30; terminal leaflet 10–40 mm. Flowers: epicalyx bractlets usually entire; hypanthium interior glabrous; sepals not red-mottled adaxially; filaments 0.5–1.5 × 0.2–0.5 mm; carpels 50–100; styles 2–3 mm. 2n = 28.

Flowering summer. Meadow edges, along streams, in chaparral; 50–1600 m; Calif.

Variety elata occurs away from the immediate coast in the northern coastal ranges (Contra Costa to Humboldt counties), western foothills of the Sierra Nevada (Butte to Tulare counties), and disjunctly in the San Bernardino Mountains. At its most distinctive, the variety is represented by relatively delicate plants with finely divided leaflets at higher elevations in the North Coast ranges. Populations in the Sierran foothills, including the type of Potentilla elata var. dissita, tend to be relatively coarse and densely villous. Intermediates with var. californica exist in interior mid elevation sites, including Mount Diablo in Contra Costa County and the type locality of Horkelia glandulosa Eastwood in Mendocino County.


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