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FOC | Family List | FOC Vol. 22 | Poaceae

86. Agrostis Linnaeus, Sp. Pl. 1: 61. 1753.

剪股颖属 jian gu ying shu

Authors: Sheng-lian Lu & Sylvia M. Phillips

Pentatherum Nábelek.

Annuals or perennials, tufted or sometimes with rhizomes or stolons. Leaf blades linear to filiform or setaceous, flat or rolled; ligule membranous. Inflorescence a panicle, open to contracted or spikelike. Spikelets with 1 floret, small, often gaping, without rachilla extension; rachilla disarticulating above glumes; glumes persistent, longer than floret, subequal or lower a little longer, membranous, 1-veined, apex subacute to acuminate; floret callus glabrous or shortly pilose; lemma oblong to elliptic, thinner than glumes, often hyaline, 5-veined, rounded on back, glabrous or hairy, lateral veins sometimes excurrent, awnless or awned from back, apex truncate or toothed; awn usually geniculate, sometimes weakly so or straight when short; palea shorter than lemma, sometimes very small. Stamens 3. Caryopsis oblong, sulcate on ventral side.

About 200 species:temperate and cold regions of the N hemisphere, also on tropical mountains; 25 species (eight endemic) in China.

The genus includes some good fodder and lawn plants.

Some species are superficially very similar, and correct identification depends on careful observation of spikelet details. It is also important to collect the basal parts to show the presence or absence of rhizomes and stolons. When the palea is long it is normally obvious, but small paleas usually adhere to the mature caryopsis and may appear to be absent. The presence or absence of awns is useful for identification, but awned species may have awnless variants, and vice versa, and the position of the awn on the lemma back can also be rather variable.


1 Lemma hairy   (2)
+ Lemma glabrous   (4)
       
2 (1) Spikelets 1.5–3 mm; palea almost equaling lemma.   1 A. munroana
+ Spikelets 3–4 mm; palea clearly shorter than lemma   (3)
       
3 (2) Palea ca. 1/3 length of lemma; ligule 2.5–6 mm.   2 A. pilosula
+ Palea ca. 2/3 length of lemma; ligule 1.5–2.5 mm.   3 A. dshungarica
       
4 (1) Palea well developed, more than 1/3 lemma length   (5)
+ Palea small, 1/3 length of lemma or less, often tiny   (10)
       
5 (4) Culms up to 20 cm tall; alpine grass in Taiwan.   17 A. fukuyamae
+ Culms up to 130 cm tall; not alpine grasses   (6)
       
6 (5) Anthers 0.5–0.8 mm; palea slightly under 1/2 lemma length.   16 A. arisan-montana
+ Anthers 0.8–1.5 mm; palea 1/2–3/4 lemma length   (7)
       
7 (6) Plant stoloniferous, rhizomes absent; panicle contracted after anthesis.   4 A. stolonifera
+ Plant tufted or rhizomatous, stolons absent; panicle open after anthesis   (8)
       
8 (7) Ligule on non-flowering shoots shorter than wide; panicle branches almost smooth; plants up to 70 cm.   5 A. capillaris
+ Ligule on non-flowering shoots as long as or longer than wide; panicle branches scabrid; plants up to 150 cm   (9)
       
9 (8) Panicle green or violet tinged, branches moderately slender, branched from near base; floret callus shortly bearded, hairs 0.2–0.5 mm.   6 A. gigantea
+ Panicle dark purple tinged brown, branches very fine, bare in lower 1/3; floret callus subglabrous.   7 A. divaricatissima
       
10 (4) Lemma awnless (occasionally a short awnlet present, included within the spikelet)   (11)
+ Lemma awned, awn flexuous or geniculate, exserted from spikelet   (19)
       
11 (10) Panicle dense, spikelike, linear or linear-oblong; culms less than 30 cm   (12)
+ Panicle open to laxly contracted, not spikelike; culms 20–100 cm   (13)
       
12 (11) Leaf blades filiform, 0.4–0.8 mm wide; spikelets ca. 2.5 mm; callus subglabrous.   8 A. mackliniae
+ Leaf blades linear, 1–3 mm wide; spikelets 2.8–4 mm; callus hairs ca. 0.2 mm.   9 A. hugoniana
       
13 (11) Culms weak, scrambling, 100–130 cm long; panicle ovate, ca. 30 cm, very delicate, branches and pedicels divaricate.   10 A. brachiata
+ Culms erect, tufted, usually less than 100 cm; panicle narrowly oblong to ovate, 3–25 cm, branches and pedicels not divaricate   (14)
       
14 (13) Spikelets usually dark purple; glumes unequal; plant often less than 30 cm tall.   11 A. nervosa
+ Spikelets usually green or grayish green, occasionally violet tinged; glumes subequal; plant (20–)40–100 cm tall   (15)
       
15 (14) Anthers 0.3–0.5(–0.7) mm; spikelets 1.3–1.8 mm (if longer, anthers small)   (16)
+ Anthers 0.5–1.5 mm; spikelets 1.8–3.2 mm   (17)
       
16 (15) Leaf blades (2.5–)4–11 mm wide; glumes subacute; lemma 3/4–9/10 spikelet length; palea 0.3–0.6 mm.   12 A. micrantha
+ Leaf blades 1–5 mm wide; glumes acuminate; lemma 2/3–3/4 spikelet length; palea less than 0.25 mm.   13 A. clavata
       
17 (15) Spikelets 2.5–3.2 mm; anthers 1–1.5 mm.   14 A. kunmingensis
+ Spikelets 1.9–2.8 mm; anthers 0.5–1 mm   (18)
       
18 (17) Panicle open, elliptic to narrowly ovate; branches bare in lower part; palea less than 0.2 lemma length.   15 A. infirma
+ Panicle usually contracted, lanceolate-oblong; branches often bearing spikelets from base; palea 0.2–0.4 lemma length.   16 A. arisan-montana
       
19 (10) Awn usually arising from middle of lemma back or above; anthers 0.5–1.2 mm   (20)
+ Awn usually arising from below middle of lemma back; anthers 1–1.6 mm   (23)
       
20 (19) Panicle open, ovate to broadly elliptic; branches widely spreading to divaricate, up to 10 cm, bare in lower part   (21)
+ Panicle laxly contracted, lanceolate; branches ascending, up to 3.5 cm, with spikelets from near base   (22)
       
21 (20) Leaf blades 2–5 mm wide; panicle elliptic, branches ascending; spikelets 1.8–2.7 mm.   18 A. sozanensis
+ Leaf blades 0.5–2 mm wide; panicle ovate, branches divaricate; spikelets 2.6–3.5 mm.   19 A. hookeriana
       
22 (20) Culms 12–20 cm tall; cauline leaf blades 1–2 mm wide; spikelets dark purple; awn arising from middle of lemma.   20 A. sinorupestris
+ Culms 30–50 cm tall; cauline leaf blades 3–5 mm wide; spikelets greenish gray; awn arising from upper 1/4–1/3 of lemma.   21 A. sinocontracta
       
23 (19) Plant with creeping surface stolons.   22 A. canina
+ Plant with subterranean rhizomes or tufted   (24)
       
24 (23) Leaf blades smooth on abaxial surface; panicle branches smooth.   23 A. flaccida
+ Leaf blades scabrid on abaxial surface; panicle branches smooth or scabrid   (25)
       
25 (24) Rhizomes present, plant loosely tufted; panicle open, at least at anthesis; callus subglabrous, hairs 0.1–0.3 mm.   24 A. vinealis
+ Rhizomes absent, plant densely tufted; panicle contracted; callus bearded, hairs 0.3–0.4 mm.   25 A. turkestanica

  • List of lower taxa


     

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