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FOC | Family List | FOC Vol. 11 | Euphorbiaceae

28. Alchornea Swartz, Prodr. 6: 98. 1788.

山麻杆属 shan ma gan shu

Authors: Huaxing Qiu & Michael G. Gilbert

Caturus Loureiro; Cladodes Loureiro; Stipellaria Bentham (1854), not Klotzsch (1848).

Trees or shrubs, dioecious or monoecious; indumentum of simple hairs. Leaves alternate; stipules small, deciduous; leaf blade simple, junction with petiole usually with 2-4 glands, mostly with 2 stipels, margin crenate or dentate; pinnately veined and shortly petiolate or palmately veined and long petiolate. Inflorescences terminal and/or axillary, branched or unbranched, male often axillary, female and bisexual terminal; bracts small, usually with 2 basal glands; flowers sometimes sessile. Male flowers fascicled; calyx closed in bud, 2-5-partite, valvate; petals absent; disk absent; stamens 4-8; filaments shortly connate; anthers 2-locular, introrse; pistillode absent. Female flowers usually 1 per bract; sepals 4-8, imbricate, sometimes 1-4-glandular; ovary 2- or 3-locular; ovules 1 per locule; styles 2 or 3, free or basally connate, simple, usually long. Fruit a capsule, 2- or 3(or 4)-locular, smooth or muricate. Seed globose, usually tuberculate.

About 50 species: tropical and subtropical regions; eight species (three endemic) in China.


1 Leaves pinnately veined, stipels absent, petiole 0.5-3 cm; male inflorescence terminal, paniculate.   1 A. rugosa
+ Leaves with 3 basal veins, stipels present at base of leaf blade, petiole 2-20 cm; male inflorescence cauliflorous or at leafless node, unbranched, or if terminal then bisexual   (2)
       
2 (1) Capsules verrucate or tuberculate   (3)
+ Capsules smooth, not verrucate or tuberculate   (4)
       
3 (2) Branchlets villous; female sepals lanceolate, 2-2.5 mm; capsule subglobose, villous, verrucate.   7 A. hunanensis
+ Branchlets densely pubescent; female sepals subovate, 3-4 mm; capsule ellipsoid, puberulent, tubercled.   8 A. tiliifolia
       
4 (2) Plants monoecious; inflorescences often bisexual   (5)
+ Plants dioecious; male inflorescence spicate; female inflorescence racemose, terminal   (6)
       
5 (4) Inflorescence branched, terminal; styles filiform, ca. 8 mm; ovary tomentulose.   5 A. androgyna
+ Inflorescence not branched, axillary; styles thick, sublanceolate, 4-5 mm; ovary puberulent.   4 A. kelungensis
       
6 (4) Male inflorescence 1.5-3.5 cm; bracts ovate, ca. 2 mm.   2 A. davidii
+ Male inflorescence more than 5 cm; bracts triangular   (7)
       
7 (6) Leaves abaxially puberulent; capsule subglobose, tomentulose.   6 A. mollis
+ Leaves abaxially puberulent only along veins; capsule strongly 3-lobed, puberulent.   3 A. trewioides

  • List of lower taxa


     

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